A classification of the bird species of South America
South
American Classification Committee
American Ornithologists' Union
(Part
4)
Part 4. Apodiformes (below)
Part 1. Struthioniformes to
Ciconiiformes (click)
Part 2. Falconiformes to Charadriiformes (click)
Part 3. Columbiformes to Caprimulgiformes (click)
Part 5. Trogoniformes to Piciformes (click)
Part 6. Suboscine Passeriformes, A (Eurylaimidae and Furnariidae)
(click)
Part 7. Suboscine Passeriformes, B (Thamnophilidae to Rhinocryptidae)
(click)
Part 8. Suboscine Passeriformes, C (Tyrannidae to Pipridae) (click)
Part 9. Oscine Passeriformes, A (Vireonidae to Sturnidae) (click)
Part 10. Oscine Passeriformes, B (Motacillidae to Emberizidae)
(click)
Part 11. Oscine Passeriformes, C (Cardinalidae to end) (click)
Hypothetical List (click)
Hybrids and Dubious Taxa (click)
Literature Cited (click)
APODIFORMES 1
APODIDAE (SWIFTS) 1a
Cypseloides cherriei Spot-fronted Swift
Cypseloides cryptus White-chinned Swift 2
Cypseloides niger Black Swift (NB) 3, 4
Cypseloides lemosi White-chested Swift 3
Cypseloides rothschildi Rothschild's Swift 2, 5
Cypseloides fumigatus Sooty Swift 3
Cypseloides senex Great Dusky Swift 3a
Streptoprocne rutila Chestnut-collared Swift 6
Streptoprocne phelpsi Tepui Swift 6
Streptoprocne zonaris White-collared Swift 6a
Streptoprocne biscutata Biscutate Swift 6a
Chaetura spinicaudus Band-rumped Swift 7, 8
Chaetura cinereiventris Gray-rumped Swift 8a
Chaetura egregia Pale-rumped Swift 9
Chaetura vauxi Vaux's Swift 10, 10a, 10b, 10c
Chaetura pelagica Chimney Swift (NB) 10a, 10c
Chaetura chapmani Chapman's Swift 10a, 10c
Chaetura viridipennis Amazonian Swift 11
Chaetura meridionalis Sick's Swift 12
Chaetura brachyura Short-tailed Swift 13
Aeronautes montivagus White-tipped Swift 13a
Aeronautes andecolus Andean Swift 13a
Tachornis furcata Pygmy Swift 14
Tachornis squamata Fork-tailed Palm-Swift 15
Panyptila cayennensis Lesser Swallow-tailed Swift
1. For continuing support
for monophyly of the Apodiformes, see Kitto & Wilson (1966),
Sibley & Ahlquist (1990), Bleiweiss et al. (1994), Johansson
et al. (2001), Livezey & Zusi (2001), Mayr (2003), Fain &
Houde (2004), and Ericson et al. (2004). The Apodiformes and Caprimulgiformes
are likely sister taxa (Cracraft et al. 2004, Ericson et al. 2006,
Livezey & Zusi 2007).
1a. Most classifications use three or more subfamily designations
within the Apodidae, e.g., for New World taxa (e.g., REFs, AOU
1998), Cypseloidinae (Cypseloides, Streptoprocne),
Chaeturinae (Chaetura), and Apodinae (Aeronautes,
Panyptila, Tachornis). <incorp Lack 1956, Orr 1963>
2. Cypseloides cryptus likely forms a superspecies with
Middle American C. storeri (REFS, Chantler 1999);
some authors (REFS) propose that C. cryptus is more
closely related to, or conspecific with, C. rothschildi/C.
fumigatus.
3. Cypseloides
niger, C. lemosi, C. rothschildi, and C.
fumigatus are considered to form a superspecies (Mayr &
Short 1970, Sibley & Monroe 1990, Chantler 1999).
3a. Cypseloides senex was placed in the genus Aerornis
by Peters (1940).
4. Cypseloides niger was formerly placed in monotypic genus Nephoecetes by
many authors (e.g., (e.g.,
Cory 1918, Peters 1940, AOU <>).
5. Cypseloides rothschildi was formerly (e.g., Zimmer 1953,
Meyer de Schauensee 1970) considered conspecific with C.
fumigatus, but see REF. Proposal
needed? Cypseloides rothschildi was formerly known as "Cypseloides
major," but that name is preoccupied by a synonym
of C. senex (Zimmer 1953).
6. Streptoprocne rutila
was placed in the genus Chaetura by Peters (1940), but
see Zimmer (1953) for the transfer to Cypseloides, a restoration
of the classification of Cory
(1918), and this was followed in most
subsequent classifications (e.g.,
Phelps & Phelps 1958a, Meyer de
Schauensee 1970, Howell & Webb 1995, Chantler 1999, Ridgely
& Greenfield 2001). Streptoprocne phelpsi was also
generally placed in Cypseloides following its original
description. However, the breeding biology and plumage pattern
of both these species indicate that they belong in Streptoprocne
(Marín and Stiles 1992); they form a superspecies (Sibley
& Monroe 1990, Chantler 1999).
6a. Fjeldså & Krabbe (1990) considered Streptoprocne
zonaris and S. biscutata to form a superspecies, but
they are evidently sympatric in southeastern Brazil.
6b. The Middle American and Andean brunneitorques subspecies
group was formerly (e.g.,
Cory 1918) considered a separate species from Streptoprocne rutila, but they were treated as conspecific by Peters
(1940).
7. Central American Chaetura fumosa was formerly
considered a subspecies of C. spinicaudus (<?>)
or of C. cinereiventris (e.g., Cory 1918, REFS),
but see Marín (2000); they constitute a superspecies.
8. The correct spelling of the species name is spinicaudus
(David & Gosselin 2002a).
8a. Meyer de Schauensee (1966) suggested that the subspecies sclateri
might deserve consideration as a separate species from Chaetura
cinereiventris.
9. Chaetura egregia was formerly (e.g., Peters 1940) considered
conspecific with C. cinereiventris, but see REFS for treating
them as separate species, a return to the classification of Cory
(1918). Sibley & Monroe (1990) considered it and C.
cinereiventris, along with West Indian C. martinica,
to form a superspecies; others (REF), however, consider C.
egregia and C. cinereiventris unlikely to
even be sister species.
10. Chaetura vauxi here includes C. andrei ("Ashy-tailed
Swift"), following Marín (1997), but see also Note
12.
10a. Mayr & Short (1970) considered Chaetura vauxi,
C. pelagica, and C. chapmani to form a superspecies.
10b. The richmondi subspecies group, which includes the
South American subspecies aphanes, was formerly (e.g.,
Peters 1940) considered a separate species from Chaetura
vauxi. <check Marin
1997>
10c. Lack (1956) suggested that Chaetura
vauxi, C. pelagica, and C. chapmani might be
conspecific, but see Wetmore (1957).
11. Chaetura viridipennis was formerly (e.g., Cory 1918,
Pinto 1937, Meyer de Schauensee 1970) considered a subspecies
of C. chapmani, but Marín (1997) provided
the rationale for treating viridipennis as a separate species.
12. Chaetura meridionalis was formerly (e.g., Cory 1918,
Pinto 1937, Meyer de Schauensee 1970, Haverschmidt & Mees
1994) considered a subspecies of C. andrei; Marín
(1997), however, showed that andrei was indistinguishable
from C. vauxi aphanes, but that meridionalis
warranted treatment as a separate species. This species is presumably
closely related to C. pelagica; the two are nearly
indistinguishable except by wing formula (Chantler 1999). Called
"Ashy-tailed Swift" by Hilty (2003).
13. Ridgely et al. (2001) treated the subspecies ocypetes
as a distinct species from Chaetura brachyura based on
morphology and (anecdotal descriptions of) voice. Proposal needed?
13a. Aeronautes montivagus and A. andecola were
formerly (e.g., Cory
1918) treated in a separate genus, Micropus, but Peters
(1940) merged this into Aeronautes, but
placed andecolus in the genus
Apus.
14. Tachornis furcata was formerly (e.g., Phelps &
Phelps 1958a, Peters 1940, Meyer de Schauensee 1970) placed in
monotypic genus Micropanyptila, or (with T. squamata)
in genus Reinarda, but see (REFS, Sibley & Monroe 1990).
Called "Pygmy Palm-Swift" in Hilty (2003) [and elsewhere?].
15. Tachornis squamata was formerly (e.g., Cory 1918, Pinto 1937, Phelps & Phelps 1958a, Meyer de Schauensee 1970)
placed in genus Reinarda, sometimes including T.
furcata, but see (REFS, Sibley & Monroe 1990). Called
"Neotropical Palm-Swift" in Hilty (2003) [and elsewhere?].
TROCHILIDAE (HUMMINGBIRDS)
Topazinae
Topaza pella Crimson Topaz 25
Topaza pyra Fiery Topaz 25
Florisuga mellivora White-necked Jacobin
Florisuga fusca Black Jacobin 22
Phaethornithinae 1, 2
Eutoxeres aquila White-tipped Sicklebill
Eutoxeres condamini Buff-tailed Sicklebill
Ramphodon naevius Saw-billed Hermit 2a
Glaucis dohrnii Hook-billed Hermit 3, 3a
Glaucis aeneus Bronzy Hermit 4, 4b
Glaucis hirsutus Rufous-breasted Hermit 4, 4b
Threnetes ruckeri Band-tailed Barbthroat 3a, 6
Threnetes leucurus Pale-tailed Barbthroat 6, 7, 7a
Threnetes niger Sooty Barbthroat 5, 6, 7, 7b
Anopetia gounellei Broad-tipped Hermit 8
Phaethornis squalidus Dusky-throated Hermit
Phaethornis rupurumii Streak-throated Hermit 9, 9a
Phaethornis longuemareus Little Hermit 9, 9b
Phaethornis idaliae Minute Hermit 9b, 9c
Phaethornis nattereri Cinnamon-throated Hermit 9c, 10
Phaethornis atrimentalis Black-throated Hermit 11
Phaethornis striigularis Stripe-throated Hermit 12
Phaethornis griseogularis Gray-chinned Hermit 12a
Phaethornis ruber Reddish Hermit 9c
Phaethornis stuarti White-browed Hermit 12b
Phaethornis subochraceus Buff-bellied Hermit
Phaethornis augusti Sooty-capped Hermit 13
Phaethornis pretrei Planalto Hermit 13, 13a
Phaethornis eurynome Scale-throated Hermit
Phaethornis anthophilus Pale-bellied Hermit
Phaethornis hispidus White-bearded Hermit
Phaethornis yaruqui White-whiskered Hermit
Phaethornis guy Green Hermit
Phaethornis syrmatophorus Tawny-bellied Hermit
Phaethornis koepckeae Koepcke's Hermit 14, 14a
Phaethornis philippii Needle-billed Hermit 14a, 15, 15a
Phaethornis bourcieri Straight-billed Hermit 15a
Phaethornis longirostris Long-billed Hermit 16, 16a, 17a
Phaethornis superciliosus Long-tailed Hermit 16, 16a
Phaethornis malaris Great-billed Hermit 16, 17b
Trochilinae 1
Doryfera ludovicae Green-fronted Lancebill 18, 18a, 18b
Doryfera johannae Blue-fronted Lancebill 18
Schistes geoffroyi Wedge-billed Hummingbird 69, 69a, 69b
Augastes scutatus Hyacinth Visorbearer 68, 68a
Augastes lumachella Hooded Visorbearer 68
Colibri delphinae Brown Violetear 22c
Colibri thalassinus Green Violetear 22a
Colibri coruscans Sparkling Violetear 22b, 22d
Colibri serrirostris White-vented Violetear
Androdon aequatorialis Tooth-billed Hummingbird 17c
Heliactin bilophus Horned Sungem 69b, 71
Heliothryx barroti Purple-crowned Fairy 69b, 70
Heliothryx auritus Black-eared Fairy 70, 70b
Polytmus guainumbi White-tailed Goldenthroat 37f
Polytmus milleri Tepui Goldenthroat 37ff, 42a
Polytmus theresiae Green-tailed Goldenthroat 37g
Avocettula recurvirostris Fiery-tailed Awlbill 24
Chrysolampis mosquitus Ruby-topaz Hummingbird 26, 26a
Anthracothorax viridigula Green-throated Mango 23
Anthracothorax prevostii Green-breasted Mango 23a
Anthracothorax nigricollis Black-throated Mango 23b
Heliangelus mavors Orange-throated Sunangel
Heliangelus amethysticollis Amethyst-throated Sunangel 55
Heliangelus strophianus Gorgeted Sunangel
Heliangelus exortis Tourmaline Sunangel
Heliangelus micraster Little Sunangel 56, 56a
Heliangelus viola Purple-throated Sunangel
Heliangelus zusii Bogota Sunangel 57
Heliangelus regalis Royal Sunangel 58
Sephanoides sephaniodes Green-backed Firecrown 54c
Sephanoides fernandensis Juan Fernandez Firecrown
Discosura popelairii Wire-crested Thorntail 29
Discosura langsdorffi Black-bellied Thorntail 29, 29a
Discosura letitiae Coppery Thorntail 29, 30
Discosura conversii Green Thorntail 29
Discosura longicaudus Racket-tailed Coquette 31
Lophornis ornatus Tufted Coquette 28
Lophornis gouldii Dot-eared Coquette 27b
Lophornis magnificus Frilled Coquette 28
Lophornis delattrei Rufous-crested Coquette
Lophornis stictolophus Spangled Coquette 28
Lophornis chalybeus Festive Coquette 28, 28a
Lophornis pavoninus Peacock Coquette 28, 28a
Phlogophilus hemileucurus Ecuadorian Piedtail
Phlogophilus harterti Peruvian Piedtail
Adelomyia melanogenys Speckled Hummingbird 47f
Anthocephala floriceps Blossomcrown 47c
Aglaiocercus kingi Long-tailed Sylph 67, 67a, 67b, 67c
Aglaiocercus coelestis Violet-tailed Sylph 67c
Aglaiocercus berlepschi Venezuelan Sylph 67a, 67c
Sappho sparganura Red-tailed Comet 62e, 62f, 66a
Polyonymus caroli Bronze-tailed Comet 66a
Taphrolesbia griseiventris Gray-bellied Comet 66, 66a
Oreotrochilus chimborazo Ecuadorian Hillstar 50
Oreotrochilus estella Andean Hillstar 51, 51a
Oreotrochilus leucopleurus White-sided Hillstar 51b
Oreotrochilus melanogaster Black-breasted Hillstar
Oreotrochilus adela Wedge-tailed Hillstar
Opisthoprora euryptera Mountain Avocetbill
Lesbia victoriae Black-tailed Trainbearer 62b, 62c, 66a
Lesbia nuna Green-tailed Trainbearer 62c, 62d
Ramphomicron dorsale Black-backed Thornbill
Ramphomicron microrhynchum Purple-backed Thornbill
Chalcostigma ruficeps Rufous-capped Thornbill 62g, 63, 64c
Chalcostigma olivaceum Olivaceous Thornbill
Chalcostigma stanleyi Blue-mantled Thornbill
Chalcostigma heteropogon Bronze-tailed Thornbill
Chalcostigma herrani Rainbow-bearded Thornbill
Oxypogon guerinii Bearded Helmetcrest 62g, 62h, 63
Oreonympha nobilis Bearded Mountaineer 62g, 63
Metallura iracunda Perija Metaltail 63, 63a
Metallura tyrianthina Tyrian Metaltail 63, 63a, 63b
Metallura williami Viridian Metaltail 63, 64, 64a
Metallura baroni Violet-throated Metaltail 64
Metallura odomae Neblina Metaltail 64, 65
Metallura theresiae Coppery Metaltail 64
Metallura eupogon Fire-throated Metaltail 64
Metallura aeneocauda Scaled Metaltail 64, 64b
Metallura phoebe Black Metaltail 64
Haplophaedia aureliae Greenish Puffleg 60c, 61a
Haplophaedia assimilis Buff-thighed Puffleg 61
Haplophaedia lugens Hoary Puffleg 61a
Eriocnemis nigrivestis Black-breasted Puffleg 58a
Eriocnemis isabellae Gorgeted Puffleg 58c
Eriocnemis vestita Glowing Puffleg 58b
Eriocnemis derbyi Black-thighed Puffleg
Eriocnemis godini Turquoise-throated Puffleg 59
Eriocnemis cupreoventris Coppery-bellied Puffleg
Eriocnemis luciani Sapphire-vented Puffleg 60, 60b
Eriocnemis mosquera Golden-breasted Puffleg
Eriocnemis glaucopoides Blue-capped Puffleg
Eriocnemis mirabilis Colorful Puffleg 60a
Eriocnemis alinae Emerald-bellied Puffleg 60a
Loddigesia mirabilis Marvelous Spatuletail 70c, 70d
Aglaeactis cupripennis Shining Sunbeam 49aaa, 49b, 49c, 49d
Aglaeactis castelnaudii White-tufted Sunbeam 49b, 49d
Aglaeactis aliciae Purple-backed Sunbeam 49d
Aglaeactis pamela Black-hooded Sunbeam 49d
Coeligena coeligena Bronzy Inca 53a
Coeligena wilsoni Brown Inca 53a, 53f
Coeligena prunellei Black Inca 53a, 53f
Coeligena torquata Collared Inca 52, 53a, 53b
Coeligena phalerata White-tailed Starfrontlet
Coeligena orina Dusky Starfrontlet 53a, 54
Coeligena bonapartei Golden-bellied Starfrontlet 53, 53a, 54
Coeligena helianthea Blue-throated Starfrontlet 53a
Coeligena lutetiae Buff-winged Starfrontlet 53a, 53d
Coeligena violifer Violet-throated Starfrontlet 53a, 53c, 53d
Coeligena iris Rainbow Starfrontlet 53a, 53e
Lafresnaya lafresnayi Mountain Velvetbreast 51c
Ensifera ensifera Sword-billed Hummingbird 53g
Pterophanes cyanopterus Great Sapphirewing 54a, 54b
Boissonneaua flavescens Buff-tailed Coronet 49aaa
Boissonneaua matthewsii Chestnut-breasted Coronet
Boissonneaua jardini Velvet-purple Coronet
Ocreatus underwoodii Booted Racket-tail 62a
Urochroa bougueri White-tailed Hillstar 49aa, 49aaa
Urosticte benjamini Purple-bibbed Whitetip 62
Urosticte ruficrissa Rufous-vented Whitetip 62
Sternoclyta cyanopectus Violet-chested Hummingbird 48e
Hylonympha macrocerca Scissor-tailed Hummingbird 48e
Heliodoxa xanthogonys Velvet-browed Brilliant
Heliodoxa gularis Pink-throated Brilliant 48a
Heliodoxa branickii Rufous-webbed Brilliant 48b
Heliodoxa schreibersii Black-throated Brilliant 48c
Heliodoxa aurescens Gould's Jewelfront 49
Heliodoxa rubinoides Fawn-breasted Brilliant 49a
Heliodoxa jacula Green-crowned Brilliant
Heliodoxa imperatrix Empress Brilliant 48d
Heliodoxa leadbeateri Violet-fronted Brilliant
Clytolaema rubricauda Brazilian Ruby 48
Patagona gigas Giant Hummingbird
Heliomaster longirostris Long-billed Starthroat 71a
Heliomaster squamosus Stripe-breasted Starthroat 71b
Heliomaster furcifer Blue-tufted Starthroat
Myrtis fanny Purple-collared Woodstar
Eulidia yarrellii Chilean Woodstar 73
Rhodopis vesper Oasis Hummingbird 71c
Thaumastura cora Peruvian Sheartail
Chaetocercus mulsant White-bellied Woodstar 74, 75
Chaetocercus bombus Little Woodstar 74
Chaetocercus heliodor Gorgeted Woodstar 74
Chaetocercus astreans Santa Marta Woodstar 74, 76
Chaetocercus berlepschi Esmeraldas Woodstar 74
Chaetocercus jourdanii Rufous-shafted Woodstar 77
Myrmia micrura Short-tailed Woodstar
Microstilbon burmeisteri Slender-tailed Woodstar 72a
Calliphlox amethystina Amethyst Woodstar
Calliphlox mitchellii Purple-throated Woodstar 72
Chlorostilbon melanorhynchus Western Emerald 33
Chlorostilbon gibsoni Red-billed Emerald 33, 33a
Chlorostilbon mellisugus Blue-tailed Emerald 33, 33b
Chlorostilbon olivaresi Chiribiquete Emerald 34
Chlorostilbon aureoventris Glittering-bellied Emerald
Chlorostilbon russatus Coppery Emerald
Chlorostilbon stenurus Narrow-tailed Emerald 35, 35a
Chlorostilbon alice Green-tailed Emerald 36
Chlorostilbon poortmani Short-tailed Emerald 35a, 36a
Chlorestes notata Blue-chinned Sapphire 32, 32a
Klais guimeti Violet-headed Hummingbird
Stephanoxis lalandi Plovercrest 27, 27a
Phaeochroa cuvierii Scaly-breasted Hummingbird 19
Campylopterus largipennis Gray-breasted Sabrewing 19a
Campylopterus hyperythrus Rufous-breasted Sabrewing 19b
Campylopterus ensipennis White-tailed Sabrewing
Campylopterus falcatus Lazuline Sabrewing
Campylopterus phainopeplus Santa Marta Sabrewing
Campylopterus villaviscensio Napo Sabrewing
Campylopterus duidae Buff-breasted Sabrewing 19a
Eupetomena macroura Swallow-tailed Hummingbird 20
Chalybura buffonii White-vented Plumeleteer 47d
Chalybura urochrysia Bronze-tailed Plumeleteer 47e
Thalurania colombica Violet-crowned Woodnymph 37, 37aa
Thalurania fannyi Green-crowned Woodnymph 37, 37a
Thalurania furcata Fork-tailed Woodnymph 37b
Thalurania watertonii Long-tailed Woodnymph
Thalurania glaucopis Violet-capped Woodnymph
Aphantochroa cirrochloris Sombre Hummingbird 21, 21a
Taphrospilus hypostictus Many-spotted Hummingbird 38
Leucochloris albicollis White-throated Hummingbird
Leucippus fallax Buffy Hummingbird
Leucippus baeri Tumbes Hummingbird
Leucippus taczanowskii Spot-throated Hummingbird 37h
Leucippus chlorocercus Olive-spotted Hummingbird 37i
Amazilia chionogaster White-bellied Hummingbird 39
Amazilia viridicauda Green-and-white Hummingbird 39
Amazilia tzacatl Rufous-tailed Hummingbird 40a
Amazilia castaneiventris Chestnut-bellied Hummingbird 40a
Amazilia amazilia Amazilia Hummingbird 41
Amazilia leucogaster Plain-bellied Emerald 42, 42a
Amazilia versicolor Versicolored Emerald 42, 42a, 43, 43a
Amazilia brevirostris White-chested Emerald 42, 42a, 44
Amazilia franciae Andean Emerald 42, 43b, 43c, 43d
Amazilia fimbriata Glittering-throated Emerald 42a, 45, 46, 46a
Amazilia lactea Sapphire-spangled Emerald 42a, 45, 45a
Amazilia amabilis Blue-chested Hummingbird 45, 45b
Amazilia rosenbergi Purple-chested Hummingbird 45
Amazilia saucerrottei Steely-vented Hummingbird 47, 47aaa
Amazilia cyanifrons Indigo-capped Hummingbird 47
Amazilia edward Snowy-bellied Hummingbird 47, 47a, 47aa
Amazilia viridigaster Green-bellied Hummingbird 40, 47, 47b
Amazilia tobaci Copper-rumped Hummingbird 47
Chrysuronia oenone Golden-tailed Sapphire
Goethalsia bella Pirre Hummingbird 47g
Goldmania violiceps Violet-capped Hummingbird
Lepidopyga coeruleogularis Sapphire-throated Hummingbird 36c
Lepidopyga lilliae Sapphire-bellied Hummingbird 37e
Lepidopyga goudoti Shining-green Hummingbird
Damophila julie Violet-bellied Hummingbird 36b
Hylocharis eliciae Blue-throated Goldentail
Hylocharis sapphirina Rufous-throated Sapphire
Hylocharis cyanus White-chinned Sapphire 37c
Hylocharis chrysura Gilded Hummingbird 37j, 37k
Hylocharis humboldtii Humboldt's Sapphire 37d
Hylocharis grayi Blue-headed Sapphire 37d
1. The monophyly of the
Trochilidae has never been questioned. The use of the subfamily
level of classification marks the major, deep division of the
lineage that is supported by many data sets (e.g., Zusi &
Bentz 1982, Sibley & Ahlquist 1990, Bleiweiss et al. 1994,
1997, Altshuler et al. 2004). However, recent genetic data (Altshuler
et al. 2004) indicate that Topaza and Florisuga
are basal to the two traditional subfamilies. Altshuler et al.
(2004) found strong support for at least four major groups within
the traditional Trochilinae, with formal recognition awaiting
additional taxon-sampling; those groups are congruent with the
groups found by Bleiweiss et al. (1997). McGuire et al. (in press)
updated the phylogeny presented in Altshuler et al. (2004). SACC proposal passed to change linear
sequence of genera.
2. Classification of the Phaethornithinae follows Hinkelmann and
Schuchmann (1997). [Sibley
& Monroe 1990 removed Ramphodon from subfamily based
on pers comms. -- trace] SACC proposal to return to classification
of Meyer de Schauensee did not pass.
2a. Genetic data (Bleiweiss et al. 2003) and morphology (Fristch
& Schuchmann 1988) confirm that Ramphodon is a member
of the Phaethornithinae, where traditionally placed, and strongly
suggest that its sister genus is Glaucis, not Eutoxeres,
as proposed by Frtisch & Schuchmann (1988). Proposal needed to move next to Glaucis in linear
sequence.
3. Glaucis dohrnii was formerly (e.g., Meyer de Schauensee
1970, Sibley & Monroe 1990) placed in genus Ramphodon,
but see [REF] for a return to the classification of Cory (1918) and Peter (1945).
3a. The genetic difference, as measured by protein electrophoresis,
between the genera Glaucis and Threnetes is actually
less than that among species within other genera of hermits (Gill
& Gerwin 1989), thus making maintenance of two separate genera
difficult to defend. A similar result was obtained from analysis
of DNA sequence divergence values (Altshuler et al. 2004). [wait for completion ongoing DNA sequence
data before making proposal?]
4. Glaucis aeneus
and G. hirsutus form a superspecies (Sibley & Monroe 1990).
4b. Glaucis
is masculine, so the correct spellings of the species names are
aeneus and hirsutus (David & Gosselin 2002b).
5. Threnetes niger
was formerly (e.g., Cory 1918) known as T. antoniae, but
see Peters (1945).
6. Threnetes ruckeri, T. leucurus,
and T. niger form a superspecies.
7. Threnetes leucurus was formerly (e.g., Peters 1945,
Meyer de Schauensee 1970) considered a separate species from T.
niger ("Sooty Barbthroat"), but Hinkelmann and
Schuchmann (1997) provided evidence that T. niger and T.
leucurus are conspecific; unfortunately, niger
has priority over leucurus, the widespread and familiar
form. SACC proposal
passed to recognize T. leucurus as a separate species from
niger because of insufficient published evidence for their
treatment as conspecific, thus returning to the traditional classification.
7a. The northwestern subspecies cervinicauda was formerly
(e.g., Cory 1918, Chapman
1926) considered a separate species from Threnetes
leucurus, but Peters (1945) and
Zimmer (1950a) treated them as conspecific.
7b. Threnetes loehkeni ("Bronze-tailed Barbthroat")
was formerly (e.g., Meyer de Schauensee 1970) considered a separate
species, but see Hinkelmann (1988a) for rationale for treating
it as a subspecies of T. niger (and also for treating Threnetes
cristalinae Ruschi, 1975, as a synonym of loehkeni).
Proposal needed?
8. Anopetia gounellei
was formerly (e.g., Peters 1945,
Meyer de Schauensee 1970) included in Phaethornis. Hinkelmann
& Schuchmann (1997) and Schuchmann (1999) resurrected monotypic
genus Anopetia for this species; see also "Threnetes
longicauda" in "Hybrids and Dubious Taxa" list.
proposal needed?
9. Phaethornis rupurumii was formerly (e.g., Meyer de Schauensee
1970) considered a subspecies of P. squalidus, but Hinkelmann
and Schuchmann (1997) provided evidence for why it should be considered
a separate species, as it was formerly (e.g., Cory 1918, Pinto 1937, Peters
1945, Phelps & Phelps 1958a).
9a. Called "Rupurumi Hermit" by Hilty (2003). proposal needed?.
9b. Sibley & Monroe (1990) considered Phaethornis longuemareus
and P. idaliae to form a superspecies; they were
considered conspecific by Peters (1945) and Zimmer (1950a).
9c. Phaethornis idaliae, P. nattereri, P. ruber,
and <> were formerly (e.g., <check Cory 1918>Pinto 1937) placed in a separate genus, Pygmornis,
but this was merged into Phaethornis by Peters (1945).
10. See Hinkelmann (1988a, b) for rationale for considering "Phaethornis
maranhaoensis," treated as a species by Meyer de Schauensee
(1970) and Vuilleumier & Mayr (1987), as a synonym (adult
male plumage) of P. nattereri.
11. Phaethornis atrimentalis was formerly (e.g., Peters
1945, Zimmer 1950a, Meyer de Schauensee 1970) considered a subspecies
of P. longuemareus, but see Hinkelmann (1990).
11a. The southern subspecies riojae was formerly (e.g., Cory 1918) considered a separate
species from Phaethornis atrimentalis, but Peters (1945) treated them as conspecific;
Zimmer (1950a) further considered riojae not distinguishable
from atrimentalis.
12. Phaethornis striigularis was formerly (e.g., Peters
1945, Zimmer 1950a, Phelps & Phelps 1958a, Meyer de Schauensee
1970) considered a subspecies of Phaethornis longuemareus,
but see Hinkelmann (1990) for a return to the classification of
Cory (1918) and Chapman (1926).
12a. The subspecies porcullae and zonura were formerly
(e.g., Peters 1945) each considered separate species from Phaethornis
griseogularis, but see Zimmer (1950a) for rationale for treatment
of all as conspecific.
12b. Peters (1945) questioned whether Phaethornis stuarti
was a species or just a subspecies of P. ruber, but they
are clearly separate species (e.g., Zimmer 1950a, Meyer de Schauensee
1970, Schuchmann et al. 1999).
13. Sibley & Monroe (1990) considered Phaethornis augusti
and P. pretrei to form a superspecies.
13a. Phaethornis pretrei was formerly (e.g., <check Cory 1918>Pinto 1937) placed in a separate
<monotypic?> genus, Anisoterus,
but this was merged into Phaethornis by Peters (1945).
14. Recently described: Weske & Terborgh (1977).
14a. Sibley & Monroe (1990) considered Phaethornis
koepckeae and P. philippii to form a superspecies.
15. [philippii as in Peters (1945), Sibley & Monroe
(1990) and Schuchmann (1999) vs. philippi, as in Meyer
de Schauensee (1970).]
15a. Phaethornis philippii and P. bourcieri were
formerly (e.g., <check Cory 1918,>Pinto
1937) placed in a separate genus, Ametrornis, but this
was merged into Phaethornis by Peters (1945)
16. Phaethornis longirostris was formerly (e.g.,
Peters 1945, Meyer de Schauensee 1970) treated as conspecific
with P. superciliosus; Hinkelmann (1996), followed
by Hinkelmann and Schuchmann (1997), provided evidence that it
should be treated as separate species, a return to the classification
of Cory (1918) <check>, namely a three-species classification: (1)
P. longirostris of Middle America and northwestern
South America; (2) P. malaris of western Amazonia,
the eastern Guianan Shield, and southeastern Brazil; and (3) P.
superciliosus (with muelleri) of the Guianan Shield
and eastern Brazil. Thus, taxa from western Amazonia formerly
(e.g., Peters 1945, Meyer de Schauensee 1970) treated as subspecies
of P. superciliosus are considered subspecies of
P. malaris. Phaethornis longirostris and P.
superciliosus form a superspecies (Schuchmann 1999). SACC proposal pending to return
to species limits of Meyer de Schauensee.
16a. Banks et al. (2002) used "Long-billed Hermit" for
longirostris and "Rusty-breasted Hermit" for
superciliosus to avoid longer names and to avoid the implication
that longirostris and superciliosus were more closely
related to each other than either is to Great-billed Hermit (P.
malaris). SACC proposal
passed to change English names from "Western Long-tailed
Hermit" to "Long-billed Hermit for P. longirostris
and from "Eastern Long-tailed Hermit" to "Long-tailed
Hermit" for P. superciliosus.
17a. Ridgely
& Greenfield (2001) considered the subspecies baroni
of western Ecuador and northwestern Peru to deserve recognition
as a separate species from Phaethornis longirostris based
on vocal and plumage differences. SACC proposal to recognize baroni as a separate
species did not pass.
17b. See Hinkelmann (1988a) for rationale for considering Phaethornis
margarettae Ruschi tentatively as a subspecies of P. malaris.
<incorp Hinkelmann 1989>
17c. Androdon was formerly (e.g., Cory 1918, Meyer de Schauensee
1970) considered a member of the Phaethornithinae, closely related
to Ramphodon, but genetic data indicate (Sibley & Ahlquist
1990, Bleiweiss et al. 1994, 1997, 2003, Altshuler et al. 2004)
that Androdon belongs in the Trochilinae and is not closely
related to Ramphodon. Schuchmann (1995) showed that this
was also consistent with behaviorial information. Schuchmann (1995)
also proposed that Androdon and Doryfera could be
placed in their own subfamily, but this is not consistent with
recent phylogenetic data (McGuire et al., in press).
18. Doryfera was formerly (e.g., Cory 1918, Peters 1945,
Meyer de Schauensee 1970) placed in the Phaethornithinae, but
see Bleiweiss et al. (1994, 1997), and Altshuler et al. (2004).
18a. The Central American subspecies veraguensis was formerly
(e.g., Cory 1918) considered
a separate species from Doryfera
ludovicae, but Peters (1945) treated
them as conspecific.
18b. The species name was given as "ludoviciae"
by Cory (1918) and Meyer de Schauensee 1966, 1970), but all indications
are that ludovicae is correct, certainly by the criterion
of prevaling usage (N. David, pers. comm.)
19. Stiles & Skutch (1989) noted that the monotypic genus
Phaeochroa perhaps could be merged into Campylopterus;
Schuchmann (1999) merged Phaeochroa into Campylopterus,
and this was followed by Dickinson (2003). SACC proposal passed to split Phaeochroa
from Campylopterus, thus restoring
the traditional classifications (e.g., Peters 1945, Meyer de Schauensee
1970) that recognize this monotypic genus.
19a. The obscurus subspecies group was formerly (e.g., Cory 1918, Chapman 1926,
Pinto 1937) considered a separate species from Campylopterus largipennis, but they were treated as conspecific by Peters
(1945).
19b. Sibley & Monroe 1990 considered
Campylopterus hyperythrus and C. duidae to
form a superspecies; they were formerly (e.g., Peters 1945) considered
conspecific; Schuchmann et al. (1999) did not consider them even
to be sister species (but no rationale provided).
20. Schuchmann (1999) merged Eupetomena into Campylopterus,
but little justification provided. SACC proposal to lump Eupetomena into Campylopterus
did not pass.
21. Schuchmann (1999) merged Aphantochroa into Campylopterus,
but little justification provided. proposal needed.
21a. Species name occasionally (e.g., Meyer de Schauensee 1970)
misspelled as "cirrhochloris."
22. Florisuga fusca was formerly (e.g., <check Cory 1918,>Pinto 1937, Peters
1945, Meyer de Schauensee 1970) placed in the monotypic genus
Melanotrochilus, which Schuchmann (1999)
merged into Florisuga, following Zimmer (1950b). proposal needed?
22a. The South American cyanotus subspecies group was formerly
(e.g., Cory 1918, Chapman
1926) treated as a separate species
from Mexican Colibri thalassinus, but they were
treated as conspecific by Peters (1945).
22b. The Tepui region subspecies germana was formerly (e.g., Cory 1918) considered a separate
species from Colibri coruscans, but Peters (1945) treated them as conspecific.
22c. SACC proposal passed
to remove hyphen in the English names of the "Violet-ears".
22d. Colibri coruscans was
formerly (e.g., Cory 1918) known as C. iolatus, but see
Peters (1945).
23. [note on superspecies?]
23a. <check>The subspecies iridescens
of western Ecuador and northwestern Peru was formerly (e.g., Zimmer
1950c) included in Anthracothorax prevostii, but
see Ridgely & Greenfield (2001) for inclusion in A. nigricollis,
as in Peters (1945); iridescens itself may deserve recognition
as a separate species Ridgely & Greenfield (2001).
23b. Anthracothorax nigricollis
was formerly (e.g., Cory 1918) known as A. violicauda,
but see Peters (1945).
24. Avocettula recurvirostris
was merged into Anthracothorax by Schuchmann (1999), pointing
out that unusual bill shape is the only difference between it
and Anthracothorax. SACC
proposal passed to recognize monotypic genus Avocettula.
25. Schuchmann (1982, 1999) considered Topaza pyra conspecific
with T. pella, but see Hu et al. (2000); the two species
are nearly parapatric in southern Venezuela with no sign of intermediacy.
SACC proposal to lump
these two species did not pass. The two
species of Topaza form a superspecies (REF).
26. Called "Ruby Topaz" in Schuchmann (1999). proposal needed?
26a. Chrysolampis
mosquitus was formerly
(e.g., Cory 1918, Pinto 1937) known as C. elatus.
27. The southern subspecies loddigesii
was formerly (e.g.,
Cory 1918, Pinto 1937) considered a separate species from Stephanoxis lalandi,
but Peters (1945) treated them as conspecific.
27a. Formerly (e.g., Meyer de Schauensee 1970) known as "Black-breasted
Plovercrest." proposal
needed?
27b. The subspecies verreauxi was formerly (e.g., Cory 1918) considered a separate
species from Lophornis chalybeus, but Peters (1945) and Zimmer (1950c) treated them
as conspecific.
28. Lophornis is masculine,
so the correct spelling of the species' names are ornatus,
magnificus, stictolophus, chalybeus, and
pavoninus (David & Gosselin 2002b).
28a. Lophornis chalybeus and L. pavoninus were formerly
(e.g., Peters 1945) placed in a separate genus, Polemistria,
but see Zimmer (1950c) for rationale for its merger into Lophornis.
29. Discosura popelairii, D. langsdorffi, D.
letitiae, and D. conversii were formerly (e.g., Peters
1945, Meyer de Schauensee 1970, Sibley & Monroe 1990) placed
in genus Popelairia, but most recent classifications
(e.g., AOU 1998, Schuchmann 1999) merge this into Discosura.
proposal needed? Zimmer (1950c) further merged Discosura
into Lophornis, and this was followed by Phelps & Phelps
(1958a).
29a. Discosura popelairii was formerly (e.g., <check Cory 1918,>
Pinto 1937) placed in the <monotypic?> genus Gouldomyia, but this was merged
into Popelairia by Peters (1945).
30. Discosura letitiae
is known from only two old specimens from Bolivia without precise
locality. Although there has been some question concerning the
validity of this taxon, Graves (1999) showed that all evidence
favors its continued treatment as a valid species.
30a. Graves (1999) proposed changing the English name of Discosura
letitiae to "Letitia's
Coquette." Proposal
needed.
31. Correct
spelling for species name is longicaudus (David & Gosselin
2002a).
32. Chlorestes notatus traditionally (e.g., Peters 1945,
Zimmer 1950c, Meyer de Schauensee 1970, Sibley & Monroe 1990)
has been treated in the monotypic genus Chlorestes;
Schuchmann (1999) merged Chlorestes into Chlorostilbon
based on morphology and voice, and this was followed by Dickinson
(2003). SACC proposal
passed to restore Chlorestes.
32a. See David &
Gosselin (2002b) for use of notata instead of notatus.
33. Species limits in the mellisugus
group of taxa in Chlorostilbon are complex. At one extreme,
Zimmer (1950d) and Schuchmann (1999) considered them all conspecific,
including the canivetii group of Middle America. Chlorostilbon
gibsoni (including nitens) was usually (e.g., Cory
1918, Peters 1945, Meyer de Schauensee 1970) considered a separate
species ("Red-billed Emerald") from C. mellisugus,
as was C. canivetii. Stiles
(1996a) proposed that C. mellisugus should be treated as
at least three separate species within South America: melanorhynchus
(of western Colombia and Ecuador), gibsoni (northern and
central Colombia, NW Venezuela), and mellisugus (rest of
South America); this represents a partial return to the classification
of Cory (1918) and was followed by Ridgely & Greenfield (2001).
Together, these taxa would form a superspecies with recently described
olivaresi. Sibley & Monroe
(1990) considered C. mellisugus to form a superspecies
with Middle American Chlorostilbon species but not with
C. gibsoni, because the two were thought to be sympatric
[are they? breeding?] in the Magdalena Valley, Colombia. SACC proposal passed to follow species
limits proposed by Stiles (1996a).
33a. The subspecies chrysogaster (= "haeberlini")
and pumilus were formerly (e.g., Cory 1918) each considered separate species
from Chlorostilbon gibsoni, but they were treated as conspecific
by Peters (1945).
33b. Chlorostilbon mellisugus
was formerly (e.g., Cory 1918, Pinto 1937, Peters 1945) known
as C. prasinus, but see Zimmer (1950d) and Meyer de Schauensee (1966).
33c. The subspecies napensis
(= "vitticeps") was formerly (e.g., Cory 1918, Peters 1945) considered a separate
species from Chlorostilbon mellisugus, but they were treated as conspecific by Meyer de Schauensee (1966, 1970), and as synonyms of C. m. phoeopygus
by Schuchmann (1999). The subspecies
peruanus was also formerly (e.g., Cory 1918) considered a separate species
from Chlorostilbon mellisugus, but Peters
(1945) treated them as conspecific.
34. Recently described: Stiles (1996a).
35. Correct spelling
for species name is stenurus (David & Gosselin 2002a).
35a. Fjeldså & Krabbe (1990)
suggested that Chlorostilbon stenurus
might best be considered a subspecies of C. poortmanni.
36. Hilty (2003) suspected that Chlorostilbon alice should
be considered a subspecies of C. stenurus.
36a. The Colombian subspecies euchloris
was formerly (e.g.,
Peters 1945) considered a separate species from Chlorostilbon poortmanni,
but Cory (1918) and
Meyer de Schauensee (1966) treated them
as conspecific, and Zimmer (1950d) questioned whether euchloris
was a diagnosable taxon. <check
Schuchmann
(1999)>
36b. The subspecies panamensis
was formerly (e.g.,
Cory 1918) considered a separate species from Damophila julie, but
Peters (1945) treated them as conspecific.
36c. The
subspecies coelina was formerly (e.g., Cory 1918) considered a separate species
from Lepidopyga coeruleogularis, but Peters (1945) treated them as conspecific.
37. Thalurania colombica and T. fannyi were
formerly (e.g., Peters 1945, Zimmer 1950d, Phelps & Phelps
1958a, Meyer de Schauensee 1966, 1970) considered conspecific
with T. furcata, but Escalante-Pliego & Peterson (1992)
showed that the three taxa are essentially parapatric with no
signs of gene flow; they constitute a superspecies (Sibley &
Monroe 1990). They had been treated as separate species from T.
furcata by Cory (1918) and AOU (1983, 1998).
37a. Ridgely & Greenfield (2001) recognized the subspecies
hypochlora of southwestern Ecuador as a separate species
from Thalurania fannyi
because its plumage differences
from other Trans-Andean/Middle American Thalurania treated
as species by Escalante-Pliego & Peterson (1992) were of similar
or greater magnitude; hypochlora had previously (e.g.,
Cory 1918, Chapman 1926) been treated as a species. SACC proposal to recognize hypochlora as
a separate species did not pass.
37aa. Called "Crowned Woodnymph" in Ridgely (1976),
AOU (1983), Hilty (1986), Ridgely & Gwynne (1989), Stiles
& Skutch (1989), Sibley & Monroe (1990), and Howell &
Webb (1995); called "Violet-crowned Woodnymph" in AOU
(1998), Rodner et al. (2000), Hilty (2003), and Jones (2003);
called "Blue-crowned Woodnymph" in Eisenmann (1955)
and Sibley & Monroe (1993). Dickinson (2003) followed Schuchmann's
(1999) evidently novel "Purple-crowned Woodnymph," and
this is why SACC started with "Purple-crowned." SACC proposal passed to change to
"Violet-crowned."
37b. The subspecies nigrofasciata, jelskii , simoni,
balzalni, and eriphile were formerly (e.g., Cory 1918, Chapman 1926, Pinto 1937) each
considered separate species from Thalurania
furcata, but Peters (1945) and Zimmer
(1950d) treated them as conspecific; Cory (1918) also treated "Thalurania
tschudii Gould" as a separate species, but Peters considered
this synonymous with T. f. jelskii.
37c. Correct spelling
for species name is cyanus (David & Gosselin 2002a).
37d. Ridgely & Greenfield (2001)
recognized the subspecies humboldtii of the Chocó
region as a separate species from Hylocharis grayi
because of plumage and habitat differences, and because the lumping
of humboldtii by Peters (1945), previously treated as a
separate species (e.g., Cory 1918, Chapman 1926), was never
justified. SACC proposal
passed to recognize humboldtii as a separate species.
37e. Meyer de Schauensee
(1966) and Sibley & Monroe (1990) suggested that Lepidopyga
lilliae might be a plumage stage of L. coeruleogularis;
Peters (1945) treated it as a subspecies of L. coeruleogularis.
37ee. The subspecies luminosa was formerly (e.g., Peters
1945) treated (with phaeochroa) as a separate species from
Lepidopyga goudoti, but they were considered conspecific
by <?> Meyer de Schauensee (1966).
37f. Polytmus guainumbi was formerly (e.g.,
Cory 1918) known as Polytmus thaumantias.
37ff. Peters (1945) placed Polytmus milleri in the monotypic
genus Waldronia; Zimmer (1950e) provided rationale for
merger of Waldronia into Polytmus, which was followed
by Meyer de Schauensee (1966) and subsequent classifications..
37g. Cory (1918) and Pinto (1937) placed Polytmus theresiae
in the monotypic genus Psilomycter; Peters (1945) placed
it in the monotypic genus Smaragdites, which predated Psilomycter;
Zimmer (1950e) provided rationale for merger of Smaragdites
into Polytmus, which was followed by Meyer de Schauensee
(1966) and subsequent classifications.
37h. Leucippus taczanowskii was formerly (e.g.,
Cory 1918) treated in a separate monotypic genus, Thaumasius,
but Peters (1940) merged this into Talaphorus, which was then merged into Leucippus by
Zimmer (1950e); this
was followed by Meyer de Schauensee (1966) and subsequent classifications.
37i. Leucippus chlorocercus was formerly (e.g.,
Cory 1918, Peters 1945) treated in a separate monotypic genus,
Talaphorus, but this was merged
into Leucippus by Zimmer
(1950e); this merger was followed by Meyer de Schauensee (1966)
and subsequent classifications.
37j. Hylocharis chrysura was formerly (e.g., Cory 1918) known
as H. ruficollis, but see Peters (1945).
37k. Called "Gilded Sapphire" in Mazar
Barnett & Pearman (2001).
38. Taphrospilus hypostictus was formerly (e.g., Peters
1945) treated in the genus Talaphorus; Zimmer (1950e) merged
Talaphorus into Polytmus. Meyer de Schauensee (1966,
1970) followed the merger of Talaphorus into Leucippus,
but resurrected the monotypic genus Taphrospilus for this
species; Schuchmann (1999) followed Zimmer (1950e) in merging
Taphrospilus into Leucippus. proposal needed.
39. Amazilia chionogaster
and A. viridicauda
were formerly (e.g., Cory 1918,
Pinto 1937, Peters 1945) placed in Leucippus, but Zimmer
(1950e) transferred these two species to Amazilia, and
this was followed by Meyer de Schauensee (1966) and most subsequent
classifications. Schuchmann (1999), however, transferred these
two species back to Leucippus. proposal needed.
40. Amazilia cupreicauda was
formerly (e.g., Peters 1945, Phelps & Phelps 1958a, Meyer
de Schauensee 1970) treated
as a subspecies of A. viridigaster; Schuchmann (1999) treated
it as a separate species, based on plumage differences; Weller
(2000) provided rationale for this change. SACC proposal passed to lump cupreicauda
and viridigaster.
40a. Schuchmann (1999) considered Amazilia tzacatl and
A. castaneiventris to form a superspecies.
41. Schuchmann (1999) treated the subspecies alticola of
southern Ecuador as a separate species from Amazilia amazilia,
representing a return to the classification of Cory (1918). Not
followed by Ridgely & Greenfield (2001). Rationale now published
by Weller (2000b), and Dickinson (2003) treated it as a separate
species. SACC proposal
to recognize alticola as a separate species did not pass. The subspecies dumerilii and leucophoea
were both also formerly (e.g.,
Cory 1918) considered separate species from Amazilia
amazilia, but Peters (1945) treated
them as conspecific.
42. Schuchmann (1999) resurrected the genus Agyrtia for
these species, a return to the classification of <check Cory 1918,>.
SACC proposal passed
to continue to recognize broad Amazilia pending publication
of data and rationale to support the fragmentation.
42a. Polytmus milleri and Amazilia leucogaster,
A. versicolor, A. brevirostris, A. fimbriata,
and A. lactea were formerly (e.g., <check Cory 1918,>Pinto 1937) placed in a separate
genus, Agyrtina; Peters (1945) merged this into Amazilia
and placed milleri in Polytmus.
43. Schuchmann (1999) treated Amazilia
rondoniae as a species separate from Amazilia versicolor
on basis of sympatry, and this was followed by Dickinson (2003).
SACC proposal passed
to continue to treat rondoniae as a subspecies of A.
versicolor.
43a. The subspecies milleri, hollandi, and nitidifrons
were each formerly (e.g.,
Cory 1918) considered separate species from Amazilia
versicolor; Peters (1945) treated
milleri and nitidifrons as subspecies of Amazilia
versicolor, but retained hollandi as a separate species;
Meyer de Schauensee <?>(1966) treated hollandi as
a subspecies of A. versicolor.
43b. The subspecies viridiceps was formerly (e.g., Cory 1918) considered a separate
species from Amazilia franciae, but Peters (1945) treated them as conspecific.
43c. Amazilia franciae was formerly (e.g., Cory 1918) treated in a separate genus, Uranomitra,
but Peters (1945) merged this into Amazilia.
43d. The Peruvian subspecies cyanocollis
was formerly (e.g.,
Cory 1918) considered a separate species from Amazilia franciae,
but Peters (1945) treated them as conspecific.
44. Amazilia brevirostris was formerly known as
A. chionopectus (e.g., Peters 1945, Meyer de Schauensee
1970), but see Schuchmann (1999), a return to the species
name of <check Cory 1918,>Pinto
(1937). proposal needed?
45. Schuchmann (1999) resurrected the
genus Polyerata for these species. SACC proposal passed to continue to recognize broad
Amazilia pending publication of data and rationale to support
the fragmentation.
45a. The subspecies bartletti
was formerly (e.g.,
Cory 1918) considered a separate species from Amazilia lactea, but
Peters (1945) treated them as conspecific.
45b. The Central American subspecies decora was formerly
(e.g., Cory 1918) considered
a separate species from Amazilia
amabilis, but Peters (1945) treated
them as conspecific, and this has been followed by some recent
authors (e.g., Schuchmann 1999, Dickinson 2003) but not by Wetmore
(1968), AOU (1983, 1998), Ridgely & Gwynne (1989) or Stiles
& Skutch (1989). See Wetmore (1968) for rationale for treating
decora as a separate species.
46. Amazilia distans
("Tachira Emerald"), formerly (e.g.,
Phelps & Phelps 1958a, Meyer de Schauensee 1970) treated as a valid species, is thought
to represent a hybrid between Amazilia fimbriata and Hylocharis
cyanus (Weller & Schuchmann 1997, Graves 1998).
46a. The subspecies maculicauda,
apicalis, tephrocephala, and fluviatilis
were formerly (e.g.,
Cory 1918, Chapman 1926, Pinto 1937) each considered separate
species from Amazilia fimbriata, but Peters (1945) treated them as conspecific.
47. Schuchmann (1999) resurrected the genus Saucerottia
for these species. SACC
proposal passed to continue to recognize broad Amazilia
pending publication of data and rationale to support the fragmentation.
47a. Amazilia edward was only recently recorded for the
first time from South America, with published photographs from
northwestern Colombia (Colorado & Pulgarín 2003); previous
reports from Ecuador (in Chapman 1926) refer to mislabeled specimens.
47aa. SACC proposal
passed to change English name to "Snowy-bellied" (as in AOU 1983, 1998, Ridgely & Gwynne 1989,
and Stiles & Skutch 1989), from "Snowy-breasted"
(as in Ridgway 1911, Eisenmann 1955, Slud 1964, Wetmore 1968,
Ridgely 1976, Sibley & Monroe 1990, Schuchmann 1999, Dickinson
2003).
47aaa. Stiles & Skutch (1989) noted that the Central American
subspecies, sophiae, differs in voice and behavior from
South American saucerrottei and probably deserves recognition
as a separate species.
47b. The Venezuelan subspecies iodura, often overlooked
completely (e.g. Peters 1945), was formerly (e.g., Cory 1918) considered a separate species
from Amazilia viridigaster; Weller (1999) treated it as a valid taxon, but
a subspecies of A. viridigaster. The taxon inculta
was also formerly (e.g.,
Cory 1918) considered a separate species from A. viridigaster, but
Peters (1945) considered it a synonym of A. v. viridigaster.
47c. The genus Anthocephala was formerly (e.g., Cory 1918)
known as Simonula, but see Peters (1945). The relationships of Anthocephala
are uncertain. Schuchmann (1999) noted possible relationships
to Adelomyia or extralimital Microchera.
47d. The subspecies intermedia
of southwestern Ecuador was formerly (e.g., Cory 1918, Chapman 1926) considered a separate
species from Chalybura buffonii, but Peters (1945) tentatively treated them as
conspecific; Meyer de Schauensee (1966) and Ridgely & Greenfield
(2001) suggested that intermedia might deserve recognition
as a separate species. The subspecies caeruleogaster of
the Eastern Andes of Colombia was also formerly (e.g., Cory 1918) considered a separate species
from Chalybura buffonii, but Peters (1945) treated them as conspecific;
Meyer de Schauensee (1966) suggested that caeruleogaster
might also deserve recognition as a separate species.
47e. The Central American subspecies isaurae and melanorrhoa
were formerly (e.g.,
Cory 1918) each considered separate species from Chalybura urochrysia,
but Peters (1945) treated them as conspecific; they intergrade
in western Panama (Eisenmann & Howell 1962).
47f. The Bolivian subspecies inornata was formerly (e.g., Cory 1918) considered a separate
species from Adelomyia melanogenys, but Peters (1945) treated them as conspecific.
47g. Called "Rufous-cheeked Hummingbird" in AOU (1983,
1998), Ridgely & Gwynne (1989), and Sibley & Monroe (1990).
SACC proposal to change
to Rufous-cheeked did not pass.
48. Clytolaema rubricauda
is sometimes (e.g. REF) placed in
the genus Heliodoxa.
48a. Heliodoxa gularis was treated in the monotypic genus
Agapetornis by Cory (1918) and Agapeta by Peters
(1945); Zimmer (1951a) provided rationale for the merger of Agapeta
into Heliodoxa, and this was followed by Meyer de Schauensee
(1966) and most subsequent classifications.
48b. Heliodoxa branickii was formerly (e.g., Cory 1918,
Peters 1945) treated in a monotypic genus, Lampraster;
Zimmer (1951a) provided rationale for the merger of Lampraster
into Heliodoxa, and this was followed by Meyer de Schauensee
(1966) and most subsequent classifications.
48c. Heliodoxa schreibersii was formerly (e.g., Cory 1918,
Pinto 1937, Peters 1945) treated in a monotypic genus, Ionolaima;
Zimmer (1951a) provided rationale for the merger of Ionolaima
into Heliodoxa, and this was followed by Meyer de Schauensee
(1966) and most subsequent classifications.
48cc. The Peruvian subspecies whitelyana was formerly (e.g., Cory 1918) considered a separate
species from Heliodoxa schreibersii, but Peters (1945) treated them as conspecific.
48d. Heliodoxa imperatrix was formerly (e.g., Cory 1918,
Peters 1945) treated in a monotypic genus, Eugenia; Zimmer
(1951a) provided rationale for the merger of Eugenia into
Heliodoxa, and this was followed by Meyer de Schauensee
(1966) and most subsequent classifications.
48e. Based on plumage and morphology, the genera Sternoclyta
and Hylonympha were considered congeneric with, and members
of, the Middle American genus Eugenes by Renner & Schuchmann
(2004). SACC proposal
to lump Sternoclyta and Hylonympha into Eugenes
did not pass.
49. Heliodoxa aurescens was formerly (e.g., Cory 1918,
Pinto 1937, Peters 1945, Meyer de Schauensee 1970) placed in
a monotypic genus, Polyplancta. Gerwin and Zink (1989)
found that Polyplancta is embedded within Heliodoxa,
as anticipated by Zimmer (1951a). Sibley & Monroe (1990) and
Schuchmann (1999) placed Polyplancta in Heliodoxa
(alluding to additional supportive but evidently unpublished morphological,
and behavioral data). Altshuler et al.'s (2004) genetic data confirm
that recognition of Polyplancta would make Heliodoxa
paraphyletic.
49a. Heliodoxa rubinoides was formerly (e.g., Cory 1918)
treated in a monotypic genus, Phaiolaima; Zimmer (1951a)
provided rationale for the merger of Phaiolaima into Heliodoxa,
and this was followed by Meyer de Schauensee (1966) and most subsequent
classifications.
49aa. Ridgely & Greenfield (2001) suggested that the subspecies
leucura of the eastern slope of the Andes might deserve
recognition as a separate species from Urochroa bougueri.
49aaa. Schuchmann et al. (2001a) proposed that plumage and vocal
similarities indicated that Urochroa, Boissonneaua,
and Agleactis formed a monophyletic group; although members
of the Brilliant group (sensu Bleiweiss et al. 1997), they are
not each others' closest relatives (McGuire et al., in press).
Schuchmann (1985) proposed that morphology and behavior indicated
a close relationship between Agleactis, Pterophanes, and
Chalcostigma, but genetic data (McGuire et al. in press)
show that they are only distantly related.
49b. Aglaeactis cupripennis caumatonotus and A.
castelnaudii castelnaudii hybridize to an uncertain degree
in depto. Cuzco, Peru (Zimmer 1951b).
49c. The southern subspecies caumatonotus was formerly
(e.g., Cory 1918) considered
a separate species from Aglaeactis
cupripennis, but Peters (1945) treated
them as conspecific.
49d. Zimmer (1951b) suggested that A. aliciae could be
treated as a subspecies of Aglaeactis cupripennis; Fjeldså
& Krabbe (1990) considered all the Aglaeactis to form
a superspecies.
50. Oreotrochilus
chimborazo was formerly (e.g., Meyer de Schauensee 1970, Fjeldså & Krabbe 1990) considered a subspecies of O. estella,
but see Sibley & Monroe (1990) and Ridgely
& Greenfield (2001)
for rationale for species rank, representing a return to the classification
of Cory (1918) and Peters (1945).
51. Peters (1945) and Schuchmann (1999) treated stolzmanni
of northern Peru as a separate species from Oreotrochilus estella,
but this has not been followed by most authors, including Ridgely
& Greenfield (2001). proposal
needed?
51a. The subspecies bolivianus was formerly (e.g., Cory 1918, Peters 1945) considered
a separate species from Oreotrochilus
estella, but Zimmer (1951a) showed
that bolivianus was not even a diagnosable taxon and considered
it a synonym of estella; this was followed by Meyer de
Schauensee (1966) and subsequent classifications.
51b. Zimmer (1951a) provided rationale for why O. leucopleurus
is best treated as subspecies of Oreotrochilus estella,
but this has not been followed by subsequent authors because of
the latitudinal overlap in their ranges.
51c. The subspecies liriope and saul were formerly
(e.g., Cory 1918) both
considered separate species from Lafresnaya
lafresnayi, but Peters (1945) treated
them as conspecific.
52. Schuchmann (1999) treated the southern inca subspecies
group as a separate species from Coeligena torquata, a
return to the classification of Cory (1918), but provided little
rationale. SACC
proposal to recognize inca as a separate species did not
pass.
53. The Venezuelan subspecies eos was formerly (e.g., Cory 1918, Peters
1945) considered a separate
species from Coeligena bonapartei, but Meyer de Schauensee (1966) treated them as
conspecific. Schuchmann (1999) returned to the earlier classifications
and treated C. eos as a separate species. SACC proposal to recognize eos
as a separate species did not pass.
53a. All species of Coeligena except for C. iris were
formerly (e.g., Cory 1918) placed in the genus Helianthea,
and C. iris was placed in a separate genus, Diphogena;
these were both merged into Coeligena by Peters (1945).
53b. The subspecies conradi,
fulgidigula, and insectivora were formerly (e.g., Cory 1918) each considered
separate species from Coeligena torquata,
but Peters (1945) treated them all as conspecific.
53c. The
northern subspecies dichroura and osculans were
formerly (e.g., Cory
1918) both considered separate species from Coeligena violifer, but Peters
(1945) treated them all as conspecific.
53d. Coeligena lutetiae and C. violifer form a superspecies
(Parker et al. 1985); Schuchmann (1999) also proposed that C.
bonapartei and C. helianthea should also be included
in this superspecies. The latter two species may interbreed to
an uncertain extent (Fjeldså & Krabbe 1990).
53e. The subspecies hesperus, aurora, and eva
were formerly (e.g.,
Cory 1918) each considered separate species from Coeligena
iris, but Peters (1945) treated
them all as conspecific.
53f. Fjeldså & Krabbe (1990) considered Coeligena wilsoni and C. prunellei to form
a superspecies.
53g. Fjeldså & Krabbe (1990)
suggested that Ensifera should probably be merged into
Coeligena.
54. Coeligena orina was formerly (e.g.,
Meyer de Schauensee 1970) treated as a separate species ("Dusky
Starfrontlet"); however, it is known from a single specimen,
and its validity as a species-level taxon is uncertain; Bleiweiss
(1988a) suggested that Coeligena orina was a subspecies of C. bonapartei,
and this was tentatively followed by Schuchmann (1999) and Fjeldså
& Krabbe (1990). Krabbe et al. (2005) reported additional
specimens and concluded that it is a valid species-level taxon.
SACC proposal passed
to recognize orina as a valid species.
54a. Pterophanes
cyanoptera was formerly
(e.g., Cory 1918) known as P. temmincki, but see Peters
(1945).
54b. Fjeldså & Krabbe (1990)
suggested that Pterophanes should probably be merged into
Coeligena.
54c. The name formerly (e.g., Cory 1918) used for the genus
Sephanoides was Eustephanus, but see Peters (1945).
55. Species limits in the Heliangelus amethysticollis complex
have been fluid. The subspecies spencei, clarisse
(= "clarissae"), and laticlavius (= "laticlavus")
were formerly (e.g.,
Cory 1918) each considered a separate species from H. amethysticollis;
Peters (1945) treated laticlavius as conspecific with H.
amethysticollis, but maintained clarisse as a separate
species, with spencei as a subspecies of H. clarisse.
Zimmer (1951b) provided rationale for why clarisse should
be considered a subspecies of H. amethysticollis and for
treatment of spencei as a separate species, and this was
followed by Meyer de Schauensee (1966, 1970). Schuchmann
(1999) returned to the classification of Peters (1945), with H.
clarisse treated as a species and spencei as a subspecies
of H. clarisse; this was followed by Hilty (2003). SACC proposal to recognize clarisse
as a separate species did not pass.
56. Zimmer (1951b) provided rationale for why Heliangelus micraster
should be treated as subspecies of H. exortis, and this
was followed by Meyer de Schauensee (1966, 1970) and Sibley &
Monroe (1990), but their ranges approach in central Ecuador without
intergradation (Bleiweiss 1992, Ridgely et al. 2001). SACC proposal to treat H. micraster
and H. exortis as conspecific did not pass.
56a. Called "Flame-throated
Sunangel" in Ridgely & Greenfield
(2001). SACC proposal
to change English name did not pass.
57. Recently described (from unique type): Graves (1993). Not
recognized as a valid species by Schuchmann (1999). SACC proposal to place this taxon
on "Hybrids and Dubious Taxa" list did not pass.
58. Recently described:
Fitzpatrick et al. (1979).
58a. Using plumage characters, Schuchmann et al. (2001b) proposed
that Eriocnemis consisted of 3 species groups ([a] E.
vestita, E. godini, E. nigrivestis; [b] E. luciani
and E. cupreoventris, with E. sapphiropygia also ranked as a species; [c] E. alinae, E.
mirabilis) and three species of uncertain affinities (E.
glaucopoides, E. mosquera, E. derbyi).
58aa. The name formerly (e.g., Cory 1918) used for the genus
Eriocnemis
was Vestipedes.
58b. Eriocnemis
is feminine, so the correct spelling of the species name is vestita
(David & Gosselin 2002b).
58c. Newly described: Cortés-Diago et al. (2007). SACC proposal passed to recognize
as a species.
59. Recognition of Eriocnemis godini as a species
is controversial (Hilty & Brown 1986, Fjeldså &
Krabbe 1990). Graves (1996) suggested that godini represents
a hybrid population (between E. vestitus and E.
<>). Proposal
needed. Also, suspected as being a subspecies
of vestita (by whom?)
according to Ridgely et al. 2001.
60. The southern subspecies sapphiropygia was formerly
(e.g., Cory 1918) considered
a separate species from Eriocnemis
luciani, but Peters (1945) and most
subsequent classifications treated them as conspecific; Schuchmann
(1999), however, treated it as a separate species from E.
luciani, and Schuchmann et al. (2001) provided evidence
for this treatment. SACC
proposal to recognize sapphiropygia as separate species
did not pass. The
subspecies catharina was also formerly (e.g., Cory 1918) considered a separate species
from Eriocnemis luciani, but Peters (1945) treated them as conspecific.
60a. Sibley & Monroe
(1990) considered Eriocnemis alinae and E. mirabilis to form a superspecies,
and Schuchmann et al. (2001b) considered them to be sister species.
Berlioz (1971), however, thought that the plumage was so unusual
that the species did not belong in Eriocnemis.
60aa. The Peruvian subspecies dybowskii
was formerly (e.g.,
Cory 1918) considered a separate species from Eriocnemis
alinae, but Peters (1945) treated
them as conspecific. For continued ranking of dybowskii
as a subspecies, see Schuchmann
et al. (2001b).
60b. The subspecies paramillo
of the Western Andes was formerly (e.g., Cory 1918) considered a separate species
from Eriocnemis vestita, but Peters (1945) treated them as conspecific.
For continued ranking of paramillo as a subspecies, see
Schuchmann et al. (2001b).
60c. Cory (1918) included Haplophaedia in Eriocnemis.
They have always been considered as sister genera because of plumage
and morphological similarities (e.g., Schuchmann et al. (2000).
61. Haplophaedia assimilis was formerly (e.g., Peters 1945,
Meyer de Schauensee 1970) considered a subspecies of H.
aureliae, but Schuchmann et al. (2000) provided rationale
for treating it as a separate species, representing a return to
the classification of Cory (1918). Proposal needed (to assess
validity of this split).
The Ecuadorian subspecies russata
was also formerly (e.g.,
Cory 1918) considered a separate species from Haplophaedia aureliae,
but Peters (1945) treated them as conspecific.
61a. Zimmer (1951b) provided rationale for treatment of H.
lugens as a subspecies of Haplophaedia aureliae, but
this has not been followed by subsequent authors; see Schuchmann
et al. (2000).
62. Cory (1918), Peters (1945), Hilty & Brown (1986), Sibley
and Monroe (1990), Schuchmann (1999), and Ridgely & Greenfield
(2001) treated eastern slope taxon ruficrissa as separate
species from Urosticte benjamini. Zimmer (1951a)
provided rationale for treating them as conspecific, and this
was followed by Meyer de Schauensee (1966, 1970), but not by Hilty
& Brown, Sibley & Monroe (1990), Schuchmann (1999), or
Ridgely & Greenfield (2001). SACC proposal passed to recognize ruficrissa
as a separate species. The Peruvian
subspecies intermedia was also formerly (e.g., Cory 1918) considered a separate species
from Urosticte
benjamini, but Peters (1945) treated them as conspecific.
62a. The two southern subspecies, annae and addae,
were formerly (e.g.,
Cory 1918) each considered a separate species from Ocreatus underwoodii, but they were all treated as conspecific by Peters
(1945). Ridgely & Greenfield (2001)
suggested that addae might deserve recognition as a separate
species. The subspecies peruana was also formerly (e.g., Cory 1918) considered a separate
species from O. underwoodii, but they were treated as conspecific
by Peters (1945); Cory (1918) used "cissiurus"
for the species name, but not only does peruana have priority
but Peters (1945) considered
"cissiurus (= cissiura)"
a synonym of peruana.
62b. The name formerly (e.g.,
Cory 1918) used for the genus Lesbia was Psalidoprymna, but see Peters (1945).
62c. The taxon eucharis was formerly
(e.g., Cory 1918) considered
a distinct species of uncertain origin; Peters
(1945) treated it as subspecies of Lesbia victoriae but
noted that it might only be a "variant" of that species;
Zimmer (1951b), however, concluded that it belongs with L.
nuna, not L. victoriae, but did not treat it as a valid
taxon. Schuchmann (1999) treated it as a Peruvian subspecies of
L. nuna, but subsequent analyses by Weller & Schuchmann
(2004) considered eucharis to most likely represent hybrids
between L. gouldii (see Note 62d) and L. victoriae
that came from somewhere in the northern Andes (not Peru). The
southern subspecies berlepschi was also formerly (e.g.,
Cory 1918) considered a separate species from L. victoriae,
but Peters (1945) treated them as conspecific.
62d. The subspecies pallidiventris of northern Peru was
formerly (e.g., Cory
1918) considered a separate species from Lesbia
nuna, but Peters (1945) treated
them as conspecific. The northern gouldii subspecies group
(including gracilis and "chlorura") was
formerly (e.g., Cory
1918) considered a separate species from Lesbia
nuna, but they were
treated as conspecific by Peters (1945). Recently, the gouldii
subspecies group was again considered a separate species from
L. nuna by Weller & Schuchmann (2004) based on plumage
and morphology. SACC
proposal to recognize gouldii as a separate species did
not pass. The form "chlorura"
was considered a synonym of Lesbia
nuna eucharis by Schuchmann
(1999), following Zimmer (1951b); see also Note 62c.
62e. Sappho sparganura was formerly
(e.g., Cory 1918) placed in the genus Lesbia, but see Peters
(1945).
62f. The southern subspecies sapho was formerly (e.g., Cory 1918) considered a separate
species from Sappho sparganura, but Peters (1945) treated them as conspecific;
see Peters (1945) and Zimmer (1952a) for the complex nomenclature
of this species.
62g. Fjeldså (1992) proposed that Oreonympha and
Oxypogon were sister taxa based on plumage, as is suggested
by their traditional placement in linear sequences. Genetic data
(McGuire et al., in press), however, indicate that although closely
related, they are nested within Chalcostigma.
62h. The subspecies lindenii, cyanolaemus, and stubelii
were formerly (e.g.,
Cory 1918) each considered separate species from Oxypogon guerinii,
but Peters (1945) treated them all as conspecific.
63. Schuchmann & Heindl (1997) and Heindl & Schuchmann
(1998) proposed that Metallura and Chalcostigma were sister
genera, based in part on the similarities in plumage and morphology
between C. ruficeps and Metallura; in fact, C.
ruficeps was placed in Metallura by Peters (1945) See
also Note 64d. . Although the sister relationship was confirmed
by genetic data (McGuire et al., in press), Chalcostigma
itself is paraphyletic with respect to Oxypogon and Oreonympha.
63a. Metallura iracunda and M. tyrianthina
form a superspecies (Heindl & Schuchmann 1998).
63b. The subspecies chloropogon
of the Coastal Range of Venezuela was formerly (e.g., Cory 1918) considered a separate species
from Metallura tyrianthina,
but Peters (1945) treated them as conspecific. The southern smaragdinicollis
subspecies group was also formerly (e.g., Cory 1918) considered a separate species
from M. tyrianthina, but they were treated as conspecific
by Peters (1945).
64. Metallura williami, M. baroni, M. odomae,
M. theresiae, M. eupogon, and M. aeneocauda
form a superspecies (Zimmer 1952a, Graves 1980, Sibley & Monroe
1990, Heindl and Schuchmann 1998), although there is some limited
geographic overlap where some species come in contact (Ridgely & Greenfield 2001). Their close relationship is supported by genetic
data (García-Moreno et al. 1999a), with the northern species
group (M. williami, M. baroni, M. odomae)
almost certainly forming a monophyletic group, and M. aeneocauda
basal within the superspecies. García-Moreno et al. (1999a),
however, found that M. phoebe was nested within this superspecies,
contrary to Heindl & Schuchmann (1998), who considered it
as the sister taxon to that superspecies. McGuire et al. (in press),
however, found that M. phoebe not only was not part of
the M. aeneocauda superspecies but was basal to all other
Metallura. Metallura baroni was formerly (e.g., Peters 1945) treated as a subspecies of distant
Metallura eupogon, but Zimmer (1952a) provided rationale
for treatment of baroni as a separate species; this was
followed by Meyer de Schauensee (1966) and subsequent classifications.
64a. The subspecies primolina and
atrigularis were formerly (e.g.,
Cory 1918) each considered a separate species from Metallura
williami, but Peters (1945) and
Zimmer (1952a) treated them as conspecific. García-Moreno
et al.'s (1999a) genetic data suggest that primolina is
more closely related to M. odomae or M. phoebe than
it