Peer-reviewed publications

If you have institutional access, the links will take you to the article on the journal's website. If not, you may request a reprint by clicking on the relevant 'pdf' link. If you don't find what you're looking for or if you have questions, please feel free to contact me. My Google Scholar profile is here.


Demos, TC, AS Achmadi, H Handika, Maharadatunkamsi, KC Rowe & JA Esselstyn. 2017. A new species of shrew (Soricomorpha: Crocidura) from Java, Indonesia: possible character displacement despite interspecific gene flow. Journal of Mammalogy. doi: 10.1093/jmammal/gyw183. Link pdf

Eldridge, RA, AS Achmadi, TC Giarla, KC Rowe & JA Esselstyn. 2017. Geographic isolation and elevational gradients promote diversification in an endemic shrew on Sulawesi. Molecular Phylogenetics and Evolution. doi: 10.1016/j.ympev.2017.09.018. Link pdf

Esselstyn, JA, CH Oliveros, MT Swanson, & BC Faircloth. 2017. Investigating difficult nodes in the placental mammal tree with expanded taxon sampling and thousands of ultraconserved elements. Genome Biology and Evolution. doi: 10.1093/gbe/evx168. Link


Cook, JA, SE Greiman, SJ Agosta, RP Anderson, BS Arbogast, et al. 2016. Transformational principles for NEON sampling of mammalian parasites and pathogens: A response to Springer and colleagues. BioScience. 66:917–919. pdf

Demos, TC, AS Achmadi, TC Giarla, H Handika, Maharadatunkamsi, KC Rowe & JA Esselstyn. 2016. Local endemism and within-island diversification of shrews illustrate the importance of speciation in building Sundaland mammal diversity. Molecular Ecology. 25:5158–5173. doi: 10.1111/mec.13820. Link pdf

Esselstyn, JA. 2016. Speciation in shrews (Crocidura) in the Philippine Archipelago. Pp. 64–65 in Heaney, LR, DS Balete & EA Rickart, The Mammals of Luzon Island: Biogeography and Natural History of a Philippine Fauna. Johns Hopkins University Press, Baltimore.

Rowe, KC, AS Achmadi & JA Esselstyn. 2016. Repeated evolution of carnivory among Indo-Australian rodents. Evolution. 70:653–665. doi: 10.1111/evo.12871. Link pdf

Rowe, KC, AS Achmadi & JA Esselstyn. 2016. A new genus and species of omnivorous rodent (Muridae: Murinae) from Sulawesi, nested within a clade of endemic carnivores. Journal of Mammalogy. 97:978–991. doi: 10.1093/jmammal/gyw029. Link pdf


Esselstyn, JA, AS Achmadi, H Handika & KC Rowe. 2015. A hog-nosed shrew rat (Rodentia: Muridae) from Sulawesi Island, Indonesia. Journal of Mammalogy. 96:895–907. doi:10.1093/jmammal/gyv093. Link pdf

Giarla, TC & JA Esselstyn. 2015. The challenges of resolving a rapid, recent radiation: empirical and simulated phylogenomics of Philippine shrews. Systematic Biology. 64:727–740. doi:10.1093/sysbio/syv029. Link pdf

Justiniano, R, JJ Schenk, DS Balete, EA Rickart, JA Esselstyn, LR Heaney & SJ Steppan. 2015. Testing diversification models of endemic Philippine forest mice (Apomys) with nuclear phylogenies across elevational gradients reveals repeated colonization of isolated mountain ranges. Journal of Biogeography. 42:51–64. doi:10.1111/jbi.12401. Link pdf

Stanley, WT, R Hutterer, TC Giarla & JA Esselstyn. 2015. Phylogeny, phylogeography, and geographic variation in the Crocidura monax (Soricidae) species complex from the montane islands of Tanzania, with descriptions of three new species. Zoological Journal of the Linnean Society. 174:185–215. doi:10.1111/zoj.12230 Link pdf


Achmadi, AS, KC Rowe & JA Esselstyn. 2014. New records of two rarely encountered, endemic rats (Rodentia: Muridae: Murinae) from Gunung Gandangdewata, West Sulawesi Province. Treubia. 41:51–60. pdf

Amar, A & JA Esselstyn. 2014. Positive association between rat abundance and breeding success of the Critically Endangered Mariana Crow Corvus kubaryi. Bird Conservation International. 24:192–200. doi:10.1017/S095927091300018X Link pdf

Brown, RM, JA Weghorst, KV Olson, MRM Duya, AJ Barley, LV Duya, M Shekelle, I Neri-Arboleda, JA Esselstyn, NJ Dominy, PS Ong, GL Moritz, A Luczon, ML Diesmos, AC Diesmos & CD Siler. 2014. Conservation genetics of the Philippine tarsier: cryptic genetic variation restructures conservation priorities in an island archipelago primate. PLoS ONE. 9:e104340. doi: 10.1371/journal.pone.0104340 Link

Esselstyn, JA, AS Achmadi & Maharadatunkamsi. 2014. A new species of shrew (Soricomorpha: Crocidura) from West Java, Indonesia. Journal of Mammalogy. 95:216–224. doi:10.1644/13-MAMM-A-215 Link pdf

Evans, B, K Zeng, JA Esselstyn, B Charlesworth & D Melnick. 2014. Reduced representation genome sequencing suggests low diversity on the sex chromosomes of tonkean macaque monkeys. Molecular Biology and Evolution. 31:2425–2440. doi:10.1093/molbev/msu197 Link pdf

Heaney, LR, DS Balete, MJ Veluz, SJ Steppan, JA Esselstyn, AW Pfeiffer & EA Rickart. 2014. Two new species of Philippine forest mice (Apomys, Muridae, Rodentia) from Lubang and Luzon islands, with a rediscription of Apomys sacobianus Johnson, 1962. Proceedings of the Biological Society of Washington. 126:395–413. doi:10.2988/0006-324X-126.4.395 Link pdf

Kang, HJ, WT Stanley, JA Esselstyn, SH Gu & R Yanigahara. 2014. Expanded host diversity and geographic distribution of hantaviruses in Sub-Saharan Africa. Journal of Virology. 88:7663–7667. doi: 10.1128/JVI.00285-14. Link pdf

Musser, GG, AS Achmadi, JA Esselstyn & KC Rowe. 2014. Bunomys torajae, new species. In GG Musser. A systematic review of Sulawesi Bunomys (Muridae, Murinae) with the description of two new species. Bulletin of the American Museum of Natural History. 392:124–139. Link

Rowe, KC, AS Achmadi & JA Esselstyn. 2014. Convergent evolution of aquatic foraging in a new genus and species (Rodentia: Muridae) from Sulawesi Island, Indonesia. Zootaxa. 3815:541–564. doi: 10.11646/zootaxa.3815.4.5. Link pdf


Achmadi, AS, JA Esselstyn, KC Rowe, I Maryanto & MT Abdullah. 2013. Phylogeny, diversity, and biogeography of Southeast Asian endemic spiny mice (Maxomys). Journal of Mammalogy. 94:1412–1423. doi:10.1644/13-MAMM-A-092.1 Link pdf Supplementary Materials

Brown, RM, CD Siler, CH Oliveros, JA Esselstyn, AC Diesmos, PA Hosner, CW Linkem, AJ Barley, JR Oaks, MB Sanguila, LJ Welton, RG Moyle, AT Peterson & AC Alcala. 2013. Evolutionary processes of diversification in a model island archipelago. Annual Review of Ecology, Evolution, and Systematics. 44:411–435. doi:10.1146/annurev-ecolsys-110411-160323 Link pdf

Esselstyn, JA, Maharadatunkamsi, AS Achmadi, CD Siler & BJ Evans. 2013. Carving out turf in a biodiversity hotspot: Multiple previously unrecognized shrew species co-occur on Java Island, Indonesia. Molecular Ecology 22:4972–4987. doi:10.1111/mec.12450 Link pdf

Oaks, JR, J Sukumaran, JA Esselstyn, CW Linkem, CD Siler, MT Holder & RM Brown. 2013. Evidence for climate-driven diversification? A caution for interpreting ABC inferences of simultaneous historical events. Evolution. 67:991–1010. doi:10.1111/j.1558-5646.2012.01840.x Link pdf

Stanley, WT, LW Robbins, JM Malekani, SG Mbalitini, DA Migurimu, JC Mukinzi, J Hulselmans, V Prévot, E Verheyen, R Hutterer, JB Doty, BP Monroe, YJ Nakazawa, Z Braden, D Carroll, JC Kerbis Peterhans, JM Bates & JA Esselstyn. 2013. A new hero emerges: another exceptional mammalian spine and its potential adaptive significance. Biology Letters. 9:20130486. doi:10.1098/rsbl.2013.0486. Link pdf
      Press coverage: Nature, BBC.


Esselstyn, JA, AS Achmadi & KC Rowe. 2012. Evolutionary novelty in a rat with no molars. Biology Letters. doi:10.1098/rsbl.2012.0574. Link pdf Supplementary Materials
      Press coverage: Nature, CBC Radio.

Esselstyn, JA, BJ Evans, JL Sedlock, FAA Khan & LR Heaney. 2012. Single-locus species delimitation: a test of the mixed Yule-coalescent model, with an empirical application to Philippine round-leaf bats. Proceedings of the Royal Society, B. doi:10.1098/rspb.2012.0705. 279:3678–3686. Link pdf Supplementary Materials


Esselstyn, JA, SP Maher & RM Brown. 2011. Species interactions during diversification and community assembly in an island radiation of shrews. PLoS ONE. 6:e21885. doi:10.1371/journal.pone.0021885. Link

Wiles, GJ, TJ O'Shea, DJ Worthington, JA Esselstyn & EW Valdez. 2011. Status and natural history of Emballonura semicaudata rotensis on Aguiguan, Mariana Islands. Acta Chiropterologica. 13:299–309. pdf


Brown, RM, CW Linkem, CD Siler, J Sukumaran, JA Esselstyn, AC Diesmos, DT Iskandar, D Bickford, BJ Evans, JA McGuire, L Grismer, J Supriatna & N Andayani. 2010. Phylogeograpy and historical demography of Polypedates leucomystax in the islands of Indonesia and the Philippines: Evidence for recent human-mediated range expansion? Molecular Phylogenetics and Evolution. 57:598–619. Link pdf

Esselstyn, JA & SM Goodman. 2010. New species of shrew (Soricidae: Crocidura) from Sibuyan Island, Philippines. Journal of Mammalogy. 91:1467–1472. Link pdf

Esselstyn, JA & CH Oliveros. 2010. Colonization of the Philippines from Taiwan: a mulitlocus test of the biogeographic and phylogenetic relationships of isolated populations of shrews. Journal of Biogeography. 37:1504–1514. Link pdf

Esselstyn, JA, CH Oliveros, RG Moyle, AT Peterson, JA McGuire & RM Brown. 2010. Integrating phylogenetic and taxonomic data illuminates complex biogeographic patterns along Huxley's modification of Wallace's Line. Journal of Biogeography. 37:2054–2066. Link pdf Supplementary Appendix (.txt)

Patou, ML, A Wilting, P Gaubert, JA Esselstyn, C Cruan, AP Jennings, J Fickel & V Geraldine. 2010. Evolutionary history of the Paradoxurus palm civets – a new model for Asian biogeography. Journal of Biogeography. 37:2077–2097. Link pdf

Siler, CD, JR Oaks, JA Esselstyn, AC Diesmos & RM Brown. 2010. Phylogeny and biogeography of Philippine bent-toed geckos (Gekkonidae: Cyrtodactylus) contradict a prevailing model of Pleistocene diversification. Molecular Phylogenetics and Evolution. 55:699–710. Link pdf

Stanley, WT & JA Esselstyn. 2010. Biogeography and diversity among montane populations of mouse shrew (Soricidae: Myosorex) in Tanzania. Biological Journal of the Linnean Society. 100:669–680. Link pdf


Esselstyn, JA, RM Timm & RM Brown. 2009. Do geological or climatic processes drive speciation in dynamic archipelagos? The tempo and mode of diversification in Southeast Asian shrews. Evolution. 63:2595–2610. Link pdf Supplementary Appendix (.pdf)

Esselstyn, JA & RM Brown. 2009. The role of repeated sea-level fluctuations in the generation of shrew (Soricidae: Crocidura) diversity in the Philippine Archipelago. Molecular Phylogenetics and Evolution. 53:171–181. Link pdf Supplementary Appendix (.xls) Supplementary Appendix (.txt)


Amar, A, F Amidon, B Arroyo, JA Esselstyn & AP Marshall. 2008. Population trends of the forest bird community on the Pacific island of Rota, Mariana Islands. Condor. 110:421–427. Link pdf

Esselstyn, JA, HJD Garcia, MG Saulog & LR Heaney. 2008. A new species of Desmalopex (Pteropodidae) from the Philippines, with a phylogenetic analysis of the Pteropodini. Journal of Mammalogy. 89:815–825. Link pdf


Esselstyn, JA. 2007. A new species of stripe-faced fruit bat (Chiroptera: Pteropodidae: Styloctenium) from the Philippines. Journal of Mammalogy. 88:951–958. Link pdf

Esselstyn, JA. 2007. Should universal guidelines be applied to taxonomic research? Biological Journal of the Linnean Society. 90:761–764. Link pdf

2006 and earlier

Esselstyn, JA, A Amar & D Janeke. 2006. Impact of post-typhoon hunting on Mariana fruit bats (Pteropus mariannus). Pacific Science. 60:531–539. Link pdf

Esselstyn, JA, P Widmann & LR Heaney. 2004. The mammals of Palawan Island, Philippines. Proceedings of the Biological Society of Washington. 117:271–302. pdf

Esselstyn, JA, GJ Wiles & A Amar. 2004. Habitat use of the Pacific sheath-tailed bat (Emballonura semicaudata) on Aguiguan, Mariana Islands. Acta Chiropterologica. 6:303–308. Link pdf

Esselstyn, JA & RC Wildman. 1997. Observations of Juga in the diet of larval Pacific giant salamanders (Dicamptodon tenebrosus). Northwestern Naturalist. 78:70–73. Link pdf

Other publications

Esselstyn, JA. 2010. At long last, an authoritative guide to the mammals of Indochina. Journal of Mammalian Evolution. 17:215–216. [Review of: Francis, CM. 2008. A guide to the mammals of Southeast Asia. Princeton University Press.]. Link pdf

Heaney, LR, ML Dolar, DS Balete, JA Esselstyn, EA Rickart & JL Sedlock. 2010. Synopsis of Philippine mammals. Link.