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</o:shapelayout></xml><![endif]--> </head> <body bgcolor="#EAF1DD" lang=EN-US link=blue vlink=purple style='tab-interval: .5in'> <div class=WordSection1> <p class=MsoNormal align=center style='text-align:center;mso-outline-level: 1;tab-stops:4.5pt'><a name="OLE_LINK2"></a><a name="OLE_LINK1"><span style='mso-bookmark:OLE_LINK2'><b style='mso-bidi-font-weight:normal'><span style='font-size:18.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></b></span></a></p> <p class=MsoNormal align=center style='text-align:center;mso-outline-level: 1;tab-stops:4.5pt'><span style='mso-bookmark:OLE_LINK1'><span style='mso-bookmark: OLE_LINK2'><b style='mso-bidi-font-weight:normal'><span style='font-size:18.0pt; font-family:Arial'>Proposal (237 and 238) to </span></b></span></span><a href="http://www.museum.lsu.edu/%7ERemsen/SACCBaseline.htm"><span style='mso-bookmark:OLE_LINK1'><span style='mso-bookmark:OLE_LINK2'><b style='mso-bidi-font-weight:normal'><span style='font-size:18.0pt;font-family: Arial;color:blue'>South American Classification Committee</span></b></span></span></a><span style='mso-bookmark:OLE_LINK2'></span><span style='mso-bookmark:OLE_LINK1'></span><b><span style='font-size:18.0pt;font-family:Arial;color:black'><o:p></o:p></span></b></p> <p class=MsoNormal align=center style='text-align:center'><b><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></b></p> <p class=MsoNormal align=center style='text-align:center'><b><span style='font-size:16.0pt;font-family:Arial;color:black'>Proposal 237: Change linear sequence of forest Tinamidae genera<o:p></o:p></span></b></p> <p class=MsoNormal align=center style='text-align:center'><b><span style='font-size:16.0pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></b></p> <p class=MsoNormal align=center style='text-align:center'><b><span style='font-size:16.0pt;font-family:Arial;color:black'>Proposal 238: Change linear sequence of&nbsp;<i>Nothocercus species</i></span></b><span style='font-size:16.0pt;font-family:Times;mso-bidi-font-family:"Times New Roman"; color:black'><o:p></o:p></span></p> <p class=MsoNormal><b><i><span style='font-size:13.5pt;font-family:Arial; color:black'><o:p>&nbsp;</o:p></span></i></b></p> <p class=MsoNormal><b><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Proposal 237</span></u></b><span style='font-size:13.5pt; font-family:Arial;color:black'>: This proposal is to change the current linear sequence of the forest Tinamidae genera (<i>Tinamus</i>,&nbsp;<i>Nothocercus</i>&nbsp;and&nbsp;<i>Crypturellus</i>) to make it consistent with recently published molecular and morphological phylogenies. Currently, the sequence is:<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='margin-left:.25in'><i><span style='font-size:13.5pt; font-family:Arial;color:black'>Tinamus<o:p></o:p></span></i></p> <p class=MsoNormal style='margin-left:.25in'><i><span style='font-size:13.5pt; font-family:Arial;color:black'>Nothocercus<o:p></o:p></span></i></p> <p class=MsoNormal style='margin-left:.25in'><i><span style='font-size:13.5pt; font-family:Arial;color:black'>Crypturellus</span></i><span style='font-size: 13.5pt;font-family:Arial;color:black'>.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'>A &quot;YES&quot; vote to this proposal would change to:<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='margin-left:.25in'><i><span style='font-size:13.5pt; font-family:Arial;color:black'>Nothocercus<o:p></o:p></span></i></p> <p class=MsoNormal style='margin-left:.25in'><i><span style='font-size:13.5pt; font-family:Arial;color:black'>Tinamus<o:p></o:p></span></i></p> <p class=MsoNormal style='margin-left:.25in'><i><span style='font-size:13.5pt; font-family:Arial;color:black'>Crypturellus<o:p></o:p></span></i></p> <p class=MsoNormal><i><span style='font-size:13.5pt;font-family:Arial; color:black'><o:p>&nbsp;</o:p></span></i></p> <p class=MsoNormal><b><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Proposal 238</span></u></b><span style='font-size:13.5pt; font-family:Arial;color:black'>: Separately, the species sequence of&nbsp;<i>Nothocercus</i>&nbsp;is as follows:<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='margin-left:.25in'><i><span style='font-size:13.5pt; font-family:Arial;color:black'>Nothocercus bonapartei<o:p></o:p></span></i></p> <p class=MsoNormal style='margin-left:.25in'><i><span style='font-size:13.5pt; font-family:Arial;color:black'>Nothocercus julius<o:p></o:p></span></i></p> <p class=MsoNormal style='margin-left:.25in'><i><span style='font-size:13.5pt; font-family:Arial;color:black'>Nothocercus nigrocapillus<o:p></o:p></span></i></p> <p class=MsoNormal><i><span style='font-size:13.5pt;font-family:Arial; color:black'><o:p>&nbsp;</o:p></span></i></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'>A &quot;YES&quot; vote to this proposal would change to:<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='margin-left:.25in'><i><span style='font-size:13.5pt; font-family:Arial;color:black'>Nothocercus julius<o:p></o:p></span></i></p> <p class=MsoNormal style='margin-left:.25in'><i><span style='font-size:13.5pt; font-family:Arial;color:black'>Nothocercus bonapartei<o:p></o:p></span></i></p> <p class=MsoNormal style='margin-left:.25in'><i><span style='font-size:13.5pt; font-family:Arial;color:black'>Nothocercus nigrocapillus<o:p></o:p></span></i></p> <p class=MsoNormal><i><span style='font-size:13.5pt;font-family:Arial; color:black'><o:p>&nbsp;</o:p></span></i></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Discussion of generic sequence in Tinamidae:</span></u><span style='font-size:13.5pt;font-family:Arial;color:black'>&nbsp;A published phylogeny based on morphological data held&nbsp;<i>Nothocercus</i>&nbsp;to be sister to all other Tinamidae; with <i>Tinamus</i>&nbsp;sister to all genera other than&nbsp;<i>Nothocercus</i>; and <i>Crypturellus</i> sister to the remaining (aridland) genera (Bertelli et al. 2002). A phylogeny based on molecular (mtDNA) data held&nbsp;<i>Nothocercus</i>&nbsp;to be sister to (<i>Tinamus</i> + <i>Crypturellus</i>), these three forest genera together forming one of two broad Tinamidae lineages: &quot;forest&quot; and &quot;aridland&quot; (Porzecanski 2003). A strict consensus phylogeny combining both data sets found <i>Tinamus</i> + <i>Crypturellus</i> (1) to be sister, with&nbsp;<i>Nothocercus</i>&nbsp;(2) and aridland Tinamidae (3) forming three Tinamidae lineages (Bertelli &amp; Porzecanski 2004). A single most optimal phylogeny using the combined data produced similar results to the molecular tree as regards the relationship between these three genera (Bertelli &amp; Porzecanski 2004). The current linear sequence is not consistent with any of these hypotheses. However, moving <i>Nothocercus</i>&nbsp;to the start of the linear sequence would be consistent with all of them.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'>No other elements of the current generic sequence on SACC appear materially inconsistent with the Bertelli &amp; Porzecanski (2004) phylogenies.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'>Species sequence in&nbsp;<i>Nothocercus:</i>&nbsp;A separate implication of Bertelli et al. (2002) and Bertelli &amp; Porzecanski (2004) is that&nbsp;<i>Nothocercus julius</i>&nbsp;is sister to (<i>N. bonapartei</i>&nbsp;+&nbsp;<i>N. nigrocapillus</i>). The latter two species share a number of morphological features not found in <i>N. julius</i>, including one structural character (absence of rounded ocelli on the thigh) and between two and seven plumage characters (Bertelli et al. 2002). The current SACC species sequence is inconsistent with such a relationship.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Note on species sequences in other genera</span></u><span style='font-size:13.5pt;font-family:Arial;color:black'>: The&nbsp;<i>Tinamus</i>&nbsp;SACC sequence is not inconsistent with the Bertelli &amp; Porzecanski (2004) phylogeny and requires no change. The&nbsp;<i>Crypturellus</i> SACC sequence is inconsistent with these phylogenies in various respects. However, <i>Crypturellus</i>&nbsp;species level relations and species limits are controversial and difficult (see e.g. Proposals 209-211). Given that the studies referred to above did not include molecular data for all distinctive taxa in the genus&nbsp;<i>Crypturellus</i>, I do not raise a proposal to deal with the species sequence. Finally, I do not consider the aridland genera here due to a lack of familiarity with them.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Recommendations</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: A &quot;YES&quot; vote to each, for the reasons given above.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>References</span></u><span style='font-size:13.5pt;font-family: Arial;color:black'>:<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='margin-left:.25in;text-indent:-.25in'><span style='font-size:13.5pt;font-family:Arial;color:black'>Bertelli S. &amp; Porzecanski A.L. 2004. Tinamou (Tinamidae) systematics: a preliminary combined analysis of morphology and molecules.&nbsp;<i>Ornitologia Neotropical</i>&nbsp;(Suppl.) 2004. (available at http://www.arches.uga.edu/~perdiz/pdf/bertelli_&amp;_porzecanski_2004.pdf)<o:p></o:p></span></p> <p class=MsoNormal style='margin-left:.25in;text-indent:-.25in'><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='margin-left:.25in;text-indent:-.25in'><span style='font-size:13.5pt;font-family:Arial;color:black'>Bertelli S., Giannini, N.P. &amp; Goloboff, P.A. 2002. A phylogeny of the tinamous (Aves: Palaeognathiformes) based on integumentary characters.&nbsp;<i>Syst. Biol</i>. 51:959-979.<o:p></o:p></span></p> <p class=MsoNormal style='margin-left:.25in;text-indent:-.25in'><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='margin-left:.25in;text-indent:-.25in'><span style='font-size:13.5pt;font-family:Arial;color:black'>*Porzecanksi, A.L. 2003. Historical biogeography of the South American aridlands: a molecular study of endemic avian taxa. Doctoral thesis. Columbia University, New York. (*Note: I have not seen this but have relied in the above discussion on trees reproduced in Bertelli &amp; Porzecanski (2004).)</span><span style='font-size:13.5pt; font-family:Times;mso-bidi-font-family:"Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal align=center style='text-align:center'><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal align=center style='text-align:center'><span style='font-size:13.5pt;font-family:Arial;color:black'>Thomas Donegan, August 2006</span><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal align=center style='text-align:center'><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family:"Times New Roman"; color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'>===========================================================================</span><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family:"Times New Roman"; color:black'><o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><b><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Proposal 237</span></u></b><span style='font-size:13.5pt; font-family:Arial;color:black'>:</span><span style='font-size:13.5pt; font-family:Times;mso-bidi-font-family:"Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Stiles</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;Morphological and genetic data both support the change, so YES.&quot;</span><span style='font-size:13.5pt; font-family:Times;mso-bidi-font-family:"Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Stotz</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;YES. Fits biogeographically as well as&nbsp;<i>Crypturellus</i>&nbsp;and&nbsp;<i>Tinamus</i>&nbsp;are basically lowland taxa, while&nbsp;<i>Nothocercus</i>&nbsp;is Andean.&quot;</span><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family:"Times New Roman"; color:black'><o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Jaramillo</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;YES - Data look solid, and nothing is obviously &quot;off&quot; with this new arrangement. The Bertelli and Porzecanski data suggest that perhaps lumping&nbsp;<i>Tinamotis</i>&nbsp;with&nbsp;<i>Eudromia</i>&nbsp;is reasonable, but this may be a question of taste.&quot;</span><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family:"Times New Roman"; color:black'><o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Pacheco</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;YES. Os resultados parecem bem comportar a posiÁ„o de<i>&nbsp;Nothocercus</i>&nbsp;no inÌcio da sequÍncia.&quot;</span><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family:"Times New Roman"; color:black'><o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'>&nbsp;<o:p></o:p></span></p> <p class=MsoNormal><b><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Proposal 238</span></u></b><span style='font-size:13.5pt; font-family:Arial;color:black'>:</span><span style='font-size:13.5pt; font-family:Times;mso-bidi-font-family:"Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Stiles</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;Although a genetic analysis would be nice, the morphological data supporting the proposal look good, so YES.&quot;</span><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family:"Times New Roman"; color:black'><o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Stotz</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;YES. Unfortunately, I am swayed mostly by my sense of these species, rather than the data presented in the proposal, which is only moderately convincing.&quot;</span><span style='font-size:13.5pt; font-family:Times;mso-bidi-font-family:"Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Pacheco</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;YES. Vejo a proposiÁ„o, baseada em dados morfolÛgicos, como um avanÁo ‡ sequÍncia tradicional, sendo portanto favor·vel a sua implementaÁ„o.&quot;</span><span style='font-size:13.5pt;font-family: Times;mso-bidi-font-family:"Times New Roman";color:black'><o:p></o:p></span></p> </div> </body> </html>