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(51) to </span></b></span></span><span style='mso-bookmark: OLE_LINK1'><span style='mso-bookmark:OLE_LINK2'></span></span><a href="http://www.museum.lsu.edu/%7ERemsen/SACCBaseline.html"><span style='mso-bookmark:OLE_LINK1'><span style='mso-bookmark:OLE_LINK2'><b style='mso-bidi-font-weight:normal'><span style='font-size:18.0pt;font-family: Arial;color:blue'>South American Classification Committee</span></b></span></span></a><span style='mso-bookmark:OLE_LINK1'><span style='mso-bookmark:OLE_LINK2'><b style='mso-bidi-font-weight:normal'><span style='font-size:18.0pt;font-family: Arial'><o:p></o:p></span></b></span></span></p> <span style='mso-bookmark:OLE_LINK2'></span><span style='mso-bookmark:OLE_LINK1'></span> <p class=MsoNormal align=center style='text-align:center'><b style='mso-bidi-font-weight: normal'><span style='font-size:16.0pt;font-family:Arial;mso-fareast-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></b></p> <p class=MsoNormal align=center style='text-align:center'><b><span style='font-size:16.0pt;font-family:Arial;mso-fareast-font-family:"Times New Roman"; color:black'>Split&nbsp;<i>Trogon mesurus&nbsp;</i>from<i> T. melanurus</i></span></b><span style='font-size:16.0pt;font-family:Times;mso-fareast-font-family:"Times New Roman"; mso-bidi-font-family:"Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal align=center style='text-align:center'><span style='font-size:13.5pt;font-family:Times;mso-fareast-font-family:"Times New Roman"; mso-bidi-font-family:"Times New Roman";color:black'>&nbsp;<o:p></o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Effect on South American CL</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: This proposal would elevate a taxon to species rank that we currently treat, by implication, as a subspecies on our baseline list.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Background</span></u><span style='font-size:13.5pt;font-family: Arial;color:black'>: For most of their recent history (e.g., Peters 1945, Meyer de Schauensee, Sibley &amp; Monroe 1990, Collar 1996), the taxon&nbsp;<i>mesurus</i>&nbsp;has been treated as a subspecies of&nbsp;<i>Trogon melanurus</i>&nbsp;(Black-tailed Trogon). They are allopatric, highly disjunct taxa:&nbsp;<i>mesurus</i>&nbsp;is endemic to W Ecuador and NW Peru; the nearest populations of&nbsp;<i>melanurus</i>&nbsp;are in Amazonia or lowland N Colombia.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><i><span style='font-size:13.5pt;font-family:Arial; color:black'>Mesurus</span></i><span style='font-size:13.5pt;font-family:Arial; color:black'>&nbsp;is evidently &quot;identical&quot; to nominate&nbsp;<i>melanurus</i>&nbsp;except its iris color is pale, not dark (see plate in Collar (1996) Ridgely &amp; Greenfield (2001). Interestingly, it shares this pale iris with parapatric&nbsp;<i>T. comptus</i>, to which it is also very similar in plumage --&nbsp;<i>comptus</i>&nbsp;differs in having a red eyering, greener upperparts, and variable white chest-band.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>New information</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: Ridgely &amp; Greenfield (2001) considered&nbsp;<i>mesurus</i>&nbsp;a separate species from&nbsp;<i>melanurus</i>&nbsp;based largely on voice, but also plumage, iris color, and disjunct distribution. They described&nbsp;<i>mesurus</i>&nbsp;song as a &quot;slow, short series of &quot;cow&quot; notes that often starts softly and builds in strength, e.g., &quot;cuh-cuh-cuh-cuh-cow-cow-cÛw-cÛw-cÛw-cÛw. <o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'>In contrast, Ridgely &amp; Greenfield (2001) described Amazonian&nbsp;<i>melanurus</i>&nbsp;as: &quot;an often long-continued series of up to 20-30 resonant &quot;cow&quot; notes that starts softly, e.g., &quot;cuh-cuh-cuh-cuh-cow-cow-cÛw-cÛw-cÛw-cÛw.&quot; Thus, songs seem to differ primarily in length. Hilty (2003) described&nbsp;<i>melanurus</i>&nbsp;song from Venezuela as: &quot;a long ser. of rather low-pitched, inflected whistles,&nbsp;<i>waaoo, wahoo</i>, ... (or&nbsp;<i>cuuh, cuuh</i>&nbsp;...), delivered much more slowly than songs of most trogons.&quot; I don't know how to interpret the differences in these two descriptions in terms of real vs. artifact of descriptions.</span><span style='font-size:13.5pt;font-family:Times; mso-bidi-font-family:"Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'>I cannot find other published descriptions of&nbsp;<i>mesurus</i>.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'>I wondered whether&nbsp;<i>mesurus</i>&nbsp;may have been allocated to the wrong species, given its parapatry with&nbsp;<i>T. comptus</i>&nbsp;and&nbsp;<i>T. massena</i>, both clearly part of this group of trogons, and its sharing of iris color with&nbsp;<i>comptus</i>. However, the descriptions of their songs in Ridgely &amp; Greenfield (2001) differ more from that of&nbsp;<i>mesurus</i>&nbsp;than&nbsp;<i>mesurus</i>&nbsp;does from geographically more distant&nbsp;<i>melanurus&nbsp;</i>(with all appropriate caveats on interpretation of qualitative descriptions).</span><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family:"Times New Roman"; color:black'><o:p></o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'><o:p><span style='text-decoration:none'>&nbsp;</span></o:p></span></u></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Analysis</span></u><span style='font-size:13.5pt;font-family:Arial; color:black'>: From the comparative standpoint, I think Bob has a good case on treating&nbsp;<i>mesurus</i>&nbsp;as a species. In coloration, it differs as much from <i>melanurus</i>&nbsp;as does&nbsp;<i>T. comptus</i>&nbsp;or&nbsp;<i>T. massena.&nbsp;</i>In song, it seems closer to&nbsp;<i>melanurus</i>, but whether that indicates sister status, retention of a song type that is basal in the group, or convergence cannot be answered without a phylogeny for the group. I suspect there is a fair chance, given the disjunct distribution with&nbsp;<i>melanurus</i>&nbsp;and the suspicious similarity and parapatric distribution with&nbsp;<i>T</i>.&nbsp;<i>comptus</i>, that&nbsp;<i>mesurus</i>&nbsp;and&nbsp;<i>melanurus</i>&nbsp;may not be sisters. For those reasons, I would favor treating&nbsp;<i>mesurus</i>&nbsp;as a species-level taxon. Another point is that&nbsp;<i>massena</i>&nbsp;and&nbsp;<i>melanurus</i>&nbsp;could be considered more similar in coloration than either is to&nbsp;<i>mesurus</i>, yet they overlap in E Panama and are clearly two species. However, is all of that sufficient evidence for a change from status quo, especially since the explicit rationale isn't really well explicated in Ridgely &amp; Greenfield?<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Recommendation</span></u><span style='font-size:13.5pt;font-family: Arial;color:black'>: Although I personally think that there is sufficient published evidence for treating&nbsp;<i>mesurus</i>&nbsp;as a species, I very reluctantly vote &quot;NO&quot; on this proposal, only because the evidence isn't clearly presented in a coherent way in print that would allow us to cite it to defend our decision. All that is needed, in my opinion, is a very short publication that presents a few sonograms, briefly describes the coloration pattern, and makes the case that the &quot;burden-of-proof&quot; should be on those who want to treat&nbsp;<i>mesurus</i>&nbsp;as a subspecies of&nbsp;<i>melanurus</i>.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Literature Cited</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>:</span><span style='font-size:13.5pt; font-family:Times;mso-bidi-font-family:"Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal style='margin-left:.5in;text-indent:-.5in'><span style='font-size:10.0pt;font-family:Arial;color:black'>COLLAR, N. 2001. Family Trogonidae (trogons). Pp. 280-479 in &quot;Handbook of the Birds of the World, Vol. 6. Mousebirds to hornbills.&quot; (J. del Hoyo et al., eds.). Lynx Edicions, Barcelona.</span><span style='font-size:13.5pt;font-family:Times; mso-bidi-font-family:"Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal style='margin-left:.5in;text-indent:-.5in'><span style='font-size:10.0pt;font-family:Arial;color:black'>HILTY, S. L. 2003. Birds of Venezuela. Princeton University Press, Princeton, New Jersey.<o:p></o:p></span></p> <p class=MsoNormal style='margin-left:.5in;text-indent:-.5in'><span style='font-size:10.0pt;font-family:Arial;color:black'>RIDGELY , R. S., AND P. J. GREENFIELD. 2001. The birds of Ecuador. Vol. II. Field guide. Cornell University Press, Ithaca, New York.<o:p></o:p></span></p> <p class=MsoNormal align=center style='text-align:center'><span style='font-size:13.5pt;font-family:Arial;mso-fareast-font-family:"Times New Roman"; color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal align=center style='text-align:center'><span style='font-size:13.5pt;font-family:Arial;mso-fareast-font-family:"Times New Roman"; color:black'>Van Remsen, August 2003&nbsp;</span><span style='font-size:13.5pt; font-family:Times;mso-fareast-font-family:"Times New Roman";mso-bidi-font-family: "Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'>&nbsp;<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'>P.S.: If the proposal does pass, then I'll work on another one on the English names of these two.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><span style='font-size:13.5pt;font-family:Arial;color:black'>===========================</span><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family:"Times New Roman"; color:black'><o:p></o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'><o:p><span style='text-decoration:none'>&nbsp;</span></o:p></span></u></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Zimmer</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;I vote &quot;yes&quot; on splitting&nbsp;<i>mesurus</i>&nbsp;from&nbsp;<i>T. melanurus</i>. Although the published rationale for the split is a bit thin, I think that the combination of different eye color (a likely isolating mechanism) and real differences in voices, along with distributions that fit an established biogeographic pattern, all argue for a split.<span style="mso-spacerun:yes">&nbsp; </span><i>T. melanurus</i>, in my experience, bucks the tendency (noted in my comments on Proposal #49) of South American trogons to exhibit greater vocal variation. To my ears, <i>melanurus</i> almost always has a measured &quot;WOWP, WOWP, WOWP...&quot; series of notes, and the only confusion comes from variation in pitch and pace of syntopic populations of&nbsp;<i>T. viridis</i>, some of which approach the sound of&nbsp;<i>melanurus</i>. Songs of&nbsp;<i>mesurus</i>&nbsp;differ in pace and pitch -- I'm&nbsp; guessing that sonograms would show noticeable differences in note shape.&quot;</span><span style='font-size:13.5pt; font-family:Times;mso-bidi-font-family:"Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'><o:p><span style='text-decoration:none'>&nbsp;</span></o:p></span></u></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Schulenberg</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;My vote: No. It would not surprise me if this proposal were valid, but I'd prefer to wait until there is more of an analysis (even a simple minded one) for us to go on.&quot;</span><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family:"Times New Roman"; color:black'><o:p></o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'><o:p><span style='text-decoration:none'>&nbsp;</span></o:p></span></u></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Stiles</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;Trogon splits. NO to all, until all the evidence is in and published. Two or three will probably prove correct, but at this time we don't have enough solid evidence to accept them.&quot;</span><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family:"Times New Roman"; color:black'><o:p></o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'><o:p><span style='text-decoration:none'>&nbsp;</span></o:p></span></u></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Jaramillo</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;NO -- more reluctantly than for proposals 49 and 50. I think that this one is more convincing for a split with current data that is available. Nevertheless I agree that even a simple note with a couple of sonograms would be important to have in the literature for acceptance of this split.&quot;</span><span style='font-size:13.5pt;font-family: Times;mso-bidi-font-family:"Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'><o:p><span style='text-decoration:none'>&nbsp;</span></o:p></span></u></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Silva</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;No. I agree that a detailed study including all populations of this taxon is needed before to propose any taxonomic change.&quot;</span><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family: "Times New Roman";color:black'><o:p></o:p></span></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'><o:p><span style='text-decoration:none'>&nbsp;</span></o:p></span></u></p> <p class=MsoNormal><u><span style='font-size:13.5pt;font-family:Arial; color:black'>Comments from Nores</span></u><span style='font-size:13.5pt; font-family:Arial;color:black'>: &quot;NO. Idem a 49.&quot;</span><span style='font-size:13.5pt;font-family:Times;mso-bidi-font-family:"Times New Roman"; color:black'><o:p></o:p></span></p> <p class=MsoNormal align=center style='text-align:center'><span style='font-size:13.5pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> </div> </body> </html>