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mso-pagination:widow-orphan; font-size:12.0pt; font-family:"Times New Roman";} </style> <![endif]--><!--[if gte mso 9]><xml> <o:shapedefaults v:ext="edit" spidmax="1027"> <o:colormru v:ext="edit" colors="#fff1e7"/> <o:colormenu v:ext="edit" fillcolor="#fff1e7"/> </o:shapedefaults></xml><![endif]--><!--[if gte mso 9]><xml> <o:shapelayout v:ext="edit"> <o:idmap v:ext="edit" data="1"/> </o:shapelayout></xml><![endif]--> </head> <body bgcolor="#FFF1E7" lang=EN-US link=blue vlink=purple style='tab-interval: .5in'> <div class=WordSection1> <p class=MsoNormal align=center style='text-align:center'><a name="OLE_LINK2"></a><a name="OLE_LINK1"><span style='mso-bookmark:OLE_LINK2'><b style='mso-bidi-font-weight: normal'><span style='font-size:16.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'><o:p>&nbsp;</o:p></span></b></span></a></p> <p class=MsoNormal align=center style='text-align:center'><span style='mso-bookmark:OLE_LINK1'><span style='mso-bookmark:OLE_LINK2'><b style='mso-bidi-font-weight:normal'><span style='font-size:18.0pt;font-family: Arial;mso-bidi-font-family:"Times New Roman"'>Proposal (558) to </span></b></span></span><span style='mso-bookmark:OLE_LINK1'><span style='mso-bookmark:OLE_LINK2'></span></span><a href="http://www.museum.lsu.edu/~Remsen/SACCBaseline.htm"><span style='mso-bookmark:OLE_LINK1'><span style='mso-bookmark:OLE_LINK2'><b style='mso-bidi-font-weight:normal'><span style='font-size:18.0pt;font-family: Arial;mso-bidi-font-family:"Times New Roman";color:blue'>South American Classification Committee</span></b></span></span></a><span style='mso-bookmark: OLE_LINK1'><span style='mso-bookmark:OLE_LINK2'><span style='font-size:18.0pt; font-family:Arial;mso-bidi-font-family:"Times New Roman"'><o:p></o:p></span></span></span></p> <span style='mso-bookmark:OLE_LINK2'></span><span style='mso-bookmark:OLE_LINK1'></span> <p class=MsoNormal><span style='font-size:16.0pt;mso-bidi-font-size:12.0pt; font-family:Arial;mso-bidi-font-family:"Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText align=center style='margin-bottom:0in;margin-bottom:.0001pt; text-align:center'><b style='mso-bidi-font-weight:normal'><span style='font-size:16.0pt;font-family:Arial;mso-bidi-font-family:"Times New Roman"'>Treat <i style='mso-bidi-font-style:normal'>Thalurania fannyi </i>and<i style='mso-bidi-font-style:normal'> Thalurania colombica</i> as conspecific<o:p></o:p></span></b></p> <p class=MsoBlockText align=center style='margin-bottom:0in;margin-bottom:.0001pt; text-align:center'><u><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt; font-family:Arial;mso-bidi-font-family:"Times New Roman"'><o:p><span style='text-decoration:none'>&nbsp;</span></o:p></span></u></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><u><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'>Proposal</span></u><span style='font-size:14.0pt;mso-bidi-font-size: 12.0pt;font-family:Arial;mso-bidi-font-family:"Times New Roman"'>:<span style="mso-spacerun:yes">&nbsp; </span>This proposal, if it passes, would result in <i style='mso-bidi-font-style:normal'>T. colombica </i>being removed from the AOU-SACC list and lumped, a prevailing treatment in publications before the 1990s.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><u><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'>Discussion</span></u><span style='font-size:14.0pt; mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family:"Times New Roman"'>: <o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'>The split of <i style='mso-bidi-font-style:normal'>fannyi</i> from <i style='mso-bidi-font-style:normal'>colombica</i> by Escalante-Pliego &amp; Peterson (1992) was based on differences in crown coloration and under a phylogenetic species concept.<span style="mso-spacerun:yes">&nbsp; </span>Recent data shows that the distribution of <i style='mso-bidi-font-style: normal'>Thalurania </i>morphotypes in Colombia is a rather more complex matter than would have been evident from materials available in the mid-1990s. As hinted by some comments in proposal <a href="SACCprop137.htm">137</a>, this proposed split, which is currently accepted by SACC, requires reconsideration.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'>Donegan (2012) recently summarised the situation of these species in Colombia, the only country in which Escalante-Pliego &amp; Peterson (1992)'s two species supposedly co-occur.<span style="mso-spacerun:yes">&nbsp; </span>[Localities mentioned in the first two sentences are from SerranÌa de San Lucas, an isolated massif north of the Central Andes.<span style="mso-spacerun:yes">&nbsp; </span><span class=SpellE>AnorÌ</span> and <span class=SpellE>Saman·</span> are in the north of the adjacent main Central Andes range.]:<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='margin-left:.25in'><span style='font-size:14.0pt; mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family:"Times New Roman"'>&quot;</span><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:ZH-CN'>&quot;<b>WOODNYMPHS </b><i>Thalurania </i></span><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family: Arial;mso-fareast-font-family:SimSun;color:black;mso-fareast-language:JA'>sp.<o:p></o:p></span></p> <p class=MsoNormal style='margin-left:.25in;background:white'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:JA'>Trapped previously by Salaman </span><i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:ZH-CN'>et al</span></i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:JA'>. (2002a) at La Punta (5) and La <span class=SpellE>Teta</span> Resort (12) and others mist-netted at Santa Cecilia (11, 2 ) where sound-recorded in the hand or on release (XC99512-13, 104416-17) and while foraging (XC104452). All males at Santa Cecilia had a spot of purple feathers slightly behind the centre of the otherwise green crown (Fig. 5).<span style="mso-spacerun:yes">&nbsp; </span>In this feature, they are intermediate between Purple-crowned Woodnymph </span><i><span style='font-size:14.0pt; mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family:SimSun; color:black;mso-fareast-language:ZH-CN'>T. colombica </span></i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:JA'>of the East Andes and Green-crowned Woodnymph </span><i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt; font-family:Arial;mso-fareast-font-family:SimSun;color:black;mso-fareast-language: ZH-CN'>T. fannyi hypochlora </span></i><span style='font-size:14.0pt; mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family:SimSun; color:black;mso-fareast-language:JA'>of the West Andes and adjacent lowlands, which were previously treated together as 'Crowned<span style="mso-spacerun:yes">&nbsp; </span>Woodnymph'.<span style="mso-spacerun:yes">&nbsp; </span>ICN specimens collected at <span class=SpellE>AnorÌ</span> similarly possess a small purple forehead spot. Escalante-Pliego &amp; Peterson (1992) noted that '<i style='mso-bidi-font-style: normal'>One to several violet feathers at the rear edge of the forecrown <span class=GramE>are</span> observed in most Panamanian specimens</i>'. To this should now be added specimens from San Lucas and the northern Central Andes (F. G. Stiles </span><i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt; font-family:Arial;mso-fareast-font-family:SimSun;color:black;mso-fareast-language: ZH-CN'>in </span></i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt; font-family:Arial;mso-fareast-font-family:SimSun;color:black;mso-fareast-language: JA'>Remsen </span><i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt; font-family:Arial;mso-fareast-font-family:SimSun;color:black;mso-fareast-language: ZH-CN'>et al</span></i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt; font-family:Arial;mso-fareast-font-family:SimSun;color:black;mso-fareast-language: JA'>. 2012, proposal 137). </span><i><span style='font-size:14.0pt;mso-bidi-font-size: 12.0pt;font-family:Arial;mso-fareast-font-family:SimSun;color:black;mso-fareast-language: ZH-CN'>T. fannyi </span></i><span style='font-size:14.0pt;mso-bidi-font-size: 12.0pt;font-family:Arial;mso-fareast-font-family:SimSun;color:black;mso-fareast-language: JA'>and </span><i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt; font-family:Arial;mso-fareast-font-family:SimSun;color:black;mso-fareast-language: ZH-CN'>T. colombica </span></i><span style='font-size:14.0pt;mso-bidi-font-size: 12.0pt;font-family:Arial;mso-fareast-font-family:SimSun;color:black;mso-fareast-language: JA'>were split largely on the basis of crown coloration in males (Escalante-Pliego &amp; Peterson 1992). In the latter study, purple-crowned birds were considered restricted to the Santa Marta Mountains and Central Andes of Colombia, with green-crowned birds in the West Andes and adjacent lowlands. We now know that purple-crowned birds occur in the East Andes (Donegan </span><i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:ZH-CN'>et al</span></i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:JA'>. 2007), with green-crowned birds in the West Andes and purple-and-green-crowned birds in the northern Central </span><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;mso-bidi-font-family:"Times New Roman";mso-fareast-language:JA'>Andes and San Lucas. At rÌo <span class=SpellE>Saman·</span>, Caldas (05∞25'39&quot;N, 75∞01'07&quot;W), purple-and-green crowned males also occur, as do males with almost no purple in the crown (M. <span class=SpellE>Slaymaker</span> </span><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family: Arial;mso-fareast-font-family:SimSun;mso-bidi-font-family:"Times New Roman"; mso-fareast-language:ZH-CN'>in litt</span><span style='font-size:14.0pt; mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family:SimSun; mso-bidi-font-family:"Times New Roman";mso-fareast-language:JA'>. 2012). Green-and-purple-crowned males from Panama are generally assigned to </span><i style='mso-bidi-font-style:normal'><span style='font-size:14.0pt;mso-bidi-font-size: 12.0pt;font-family:Arial;mso-fareast-font-family:SimSun;mso-bidi-font-family: "Times New Roman";mso-fareast-language:ZH-CN'>T. f. fannyi</span></i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;mso-bidi-font-family:"Times New Roman";mso-fareast-language:JA'>, with pure green-crowned </span><i style='mso-bidi-font-style:normal'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;mso-bidi-font-family:"Times New Roman";mso-fareast-language:ZH-CN'>hypochlora</span></i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;mso-bidi-font-family:"Times New Roman";mso-fareast-language:ZH-CN'> </span><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;mso-bidi-font-family:"Times New Roman";mso-fareast-language:JA'>in the Colombian ChocÛ and West Andes. Treatment of </span><i style='mso-bidi-font-style: normal'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family: Arial;mso-fareast-font-family:SimSun;mso-bidi-font-family:"Times New Roman"; mso-fareast-language:ZH-CN'>T. colombica</span></i><span style='font-size:14.0pt; mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family:SimSun; mso-bidi-font-family:"Times New Roman";mso-fareast-language:ZH-CN'> </span><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;mso-bidi-font-family:"Times New Roman";mso-fareast-language:JA'>and </span><i style='mso-bidi-font-style:normal'><span style='font-size:14.0pt;mso-bidi-font-size: 12.0pt;font-family:Arial;mso-fareast-font-family:SimSun;mso-bidi-font-family: "Times New Roman";mso-fareast-language:ZH-CN'>T. fannyi</span></i><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;mso-bidi-font-family:"Times New Roman";mso-fareast-language:ZH-CN'> </span><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;mso-bidi-font-family:"Times New Roman";mso-fareast-language:JA'>as separate species requires revision in light of the known distribution of morphotypes in Colombia, identical female plumages and similar vocalisations throughout the Colombian Andes.&quot;</span><span style='font-size:14.0pt; mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family:SimSun; color:black;mso-fareast-language:JA'><o:p></o:p></span></p> <p class=MsoNormal style='background:white'><span style='font-size:14.0pt; mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family:SimSun; color:black;mso-fareast-language:JA'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'>We have more recently seen the description (recently rejected by this committee) of <i style='mso-bidi-font-style:normal'>Thalurania nigricapilla </i>(see proposal <a href="SACCprop472.htm">472</a>).<span style="mso-spacerun:yes">&nbsp; </span>The main distinguishing feature cited in the diagnosis section for the <i style='mso-bidi-font-style:normal'>nigricapilla</i> description was also in crown coloration.<span style="mso-spacerun:yes">&nbsp; </span>Most committee members accepted the proposition in the proposal that individual variation in this feature should be investigated further before <span class=SpellE>recognising</span> this species. <o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'>These two proposed woodnymph species' calls sound pretty similar (click on below links, then on &quot;Sonograms&quot;, if these do not immediately come up on your browser):<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'>http://www.xeno-canto.org/browse.php?query=Green-crowned Woodnymph (Thalurania fannyi) 8&amp;pagenumber=&amp;order=<span class=SpellE>taxonomy&amp;view</span>=3<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'>http://www.xeno-canto.org/browse.php?query=Violet-crowned Woodnymph (Thalurania colombica) 9&amp;pagenumber=&amp;order=<span class=SpellE>taxonomy&amp;view</span>=3<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'>Females are indistinguishable.<span style="mso-spacerun:yes">&nbsp; </span>In biometrics and plumage features other than adult male crown coloration, they are also similar, based on my data and experience from mist-netting populations in all three Andean cordilleras of Colombia.<span style="mso-spacerun:yes">&nbsp; </span>Mostly green-crowned birds now seem to be replaced by purple-crowned birds at some point in the mid-Central Andes (rather than this being between-Cordillera variation).<span style="mso-spacerun:yes">&nbsp; </span>Some &quot;good&quot; species show North/South distribution splits in the Central Andes, but they tend to be of higher elevation than <i style='mso-bidi-font-style:normal'>Thalurania</i>.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'>It bears note in terms of assessing this proposal that a</span><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:JA'>lthough Escalante-Pliego &amp; Peterson (1992) was published some time ago, many birders active in Colombia were unaware of the split until relatively recently, due to near-universal use of Hilty &amp; Brown (1986)'s field guide until the mid-2000s at least.<span style="mso-spacerun:yes">&nbsp; </span>The Restall and McMullan field guides split these, but they were published in 2006 and 2010/2011 respectively.<span style="mso-spacerun:yes">&nbsp; </span>The <i style='mso-bidi-font-style:normal'>status quo</i> versus novel nature of a split treatment is therefore somewhat moot in the country where the two occur together.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'>Now arguing against a change in treatment, one could adopt a similar viewpoint here to that of some committee members in proposal <a href="SACCprop173.htm">173</a> on <i style='mso-bidi-font-style:normal'>Zimmerius chrysops</i>.<span style="mso-spacerun:yes">&nbsp; </span>There, new distributional data were thought by some not to be sufficient to show current species-level taxonomy to be misled, but just to demonstrate that previous assumptions as to the distributions of species required reconsidering.<span style="mso-spacerun:yes">&nbsp; </span></span><span style='font-size:14.0pt; mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family:SimSun; color:black;mso-fareast-language:JA'>An approach against lumping these two species would regard the SACC baseline as only to be amended following a detailed peer-reviewed publication including a detailed vocal and/or molecular study specifically on topic, refuting the current treatment.<span style="mso-spacerun:yes">&nbsp; </span>No such study post- Escalante-Pliego &amp; Peterson (1992) is available and a detailed molecular and vocal investigation would indeed be helpful and welcomed.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:JA'>Although they were described very close together in time, the name <i style='mso-bidi-font-style:normal'>colombica </i>(Bourcier, 1843) apparently has priority over the name <i style='mso-bidi-font-style: normal'>fannyi </i>(Delattre &amp; Bourcier 1846).<span style="mso-spacerun:yes">&nbsp; </span>The vernacular name of this hummingbird if lumped would revert to &quot;Crowned Woodnymph&quot;.<span style="mso-spacerun:yes">&nbsp; </span>Vernacular names for this group were previously discussed in Proposal 303 (</span><a href="http://www.museum.lsu.edu/~Remsen/SACCprop303.html"><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;mso-fareast-language:JA'>http://www.museum.lsu.edu/~Remsen/SACCprop303.html</span></a><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:JA'>).<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:JA'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:JA'>Finally, I'd like to stress that Escalante-Pliego &amp; Peterson (1992)'s study includes a lot of extremely valuable data and analysis on this genus, especially as regards plumage variation and the Central American populations.<span style="mso-spacerun:yes">&nbsp; </span>The doubts expressed here as to their proposed treatment for the Colombian taxa are based in part on recent data not available to these authors and perhaps in part on a different approach to species limits.<span style="mso-spacerun:yes">&nbsp; </span>This proposal should therefore not be taken as a criticism of their work overall, which is a valuable and appreciated contribution to the ornithological literature.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-fareast-font-family: SimSun;color:black;mso-fareast-language:JA'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><u><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family: "Times New Roman"'>References</span></u><span style='font-size:14.0pt; mso-bidi-font-size:12.0pt;font-family:Arial;mso-bidi-font-family:"Times New Roman"'>:<o:p></o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:13.7pt;margin-bottom:.0001pt;text-align:justify;text-indent: -13.7pt;tab-stops:13.5pt'><span style='font-size:14.0pt;mso-bidi-font-size: 12.0pt;font-family:Arial;mso-bidi-font-family:"Times New Roman"'>Donegan, T.M. 2012.<span style="mso-spacerun:yes">&nbsp; </span>Range extensions and other notes on the birds and conservation of the SerranÌa de San Lucas, an isolated mountain range in northern Colombia.<span style="mso-spacerun:yes">&nbsp; </span><i style='mso-bidi-font-style:normal'>Bull Brit. Orn. Cl.</i><span style="mso-spacerun:yes">&nbsp; </span>132: 140-161.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:13.5pt;margin-bottom:.0001pt;text-align:justify;text-indent: -13.5pt;tab-stops:13.5pt'><span style='font-size:14.0pt;mso-bidi-font-size: 12.0pt;font-family:Arial;mso-bidi-font-family:"Times New Roman"'>Escalante-Pliego, P. &amp; Peterson, A. T. 1992. Geographic variation and species limits in Middle American woodnymphs (<i style='mso-bidi-font-style:normal'>Thalurania</i>). Wilson Bull. 104: 205&#8211;219.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:13.5pt;margin-bottom:.0001pt;text-align:justify;text-indent: -13.5pt;tab-stops:13.5pt'><span style='font-size:14.0pt;mso-bidi-font-size: 12.0pt;font-family:Arial;mso-bidi-font-family:"Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:13.5pt;margin-bottom:.0001pt;text-align:justify;text-indent: -13.5pt;tab-stops:13.5pt'><span style='font-size:14.0pt;mso-bidi-font-size: 12.0pt;font-family:Arial;mso-bidi-font-family:"Times New Roman"'>Other papers mentioned are cited in the above.<o:p></o:p></span></p> <p class=MsoBlockText align=center style='margin-bottom:0in;margin-bottom:.0001pt; text-align:center'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt; font-family:Arial;mso-bidi-font-family:"Times New Roman"'>Thomas Donegan, October 2012<o:p></o:p></span></p> <div style='mso-element:para-border-div;border:none;border-bottom:double windowtext 2.25pt; padding:0in 0in 1.0pt 0in'> <p class=MsoNormal align=center style='text-align:center;border:none; mso-border-bottom-alt:double windowtext 2.25pt;padding:0in;mso-padding-alt: 0in 0in 1.0pt 0in'><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt; font-family:Arial'><o:p>&nbsp;</o:p></span></p> </div> <p class=MsoNormal><u><span style='font-size:14.0pt;mso-bidi-font-size:19.0pt; font-family:Arial;mso-bidi-font-family:"Times New Roman"'><o:p><span style='text-decoration:none'>&nbsp;</span></o:p></span></u></p> <p class=MsoNoSpacing><u><span lang=ES style='font-size:12.0pt;mso-bidi-font-size: 19.0pt;font-family:Arial;mso-bidi-font-family:Helvetica'>Comments from Stiles</span></u><span lang=ES style='font-size:12.0pt;mso-bidi-font-size:19.0pt;font-family:Arial; mso-bidi-font-family:Helvetica'>:  </span><span style='font-size:12.0pt; mso-bidi-font-size:11.0pt;font-family:Arial;mso-bidi-font-family:"Times New Roman"; mso-ansi-language:EN-US'>YES. Although I haven t really gotten serious (yet?) about putting all my <i style='mso-bidi-font-style:normal'>Thalurania </i>data together, I am firmly convinced that there is no solid basis for splitting <i style='mso-bidi-font-style:normal'>fannyi </i>from <i style='mso-bidi-font-style: normal'>colombica.</i> To begin with, I should note that the original separation by Peterson et al. involved a bit of<span style="mso-spacerun:yes">&nbsp; </span>circular reasoning:<span style="mso-spacerun:yes">&nbsp; </span>they started out with the question of whether the green- and purple-crowned male birds were separable in multivariate space, then concluded from a PCA that they were - but mostly on the basis of crown color!<span style="mso-spacerun:yes">&nbsp; </span>They effectively negated Zimmer s statement that blue feathering at the rear of the crown in<i style='mso-bidi-font-style:normal'> fannyi </i>decreased southwards from the zone of closest approach to <i style='mso-bidi-font-style:normal'>colombica</i>; in this I find that they were correct.<span style="mso-spacerun:yes">&nbsp; </span>However, the small blue feathers bordering the crown are not the feathers of interest here: Thomas (and I) refer to the larger, more brilliant display feathers of the crown itself, and among these occurs the mixture of green and purple feathers in the N end of the Cordillera Central (Antioquia; AndrÈs Cuervo specimens) and the SerranÌa de San Lucas (BolÌvar); males of the Magdalena Valley and the Cordillera Oriental are purple-crowned.<span style="mso-spacerun:yes">&nbsp; </span>Thus, a considerable zone of intermediacy exists between <i style='mso-bidi-font-style:normal'>colombica </i>and <i style='mso-bidi-font-style:normal'>fannyi</i> based on male crown color.<span style="mso-spacerun:yes">&nbsp; </span>Even more convincing is the situation in females, in which geographic trends in coloration exist but do not support a split along the lines of male crown color.<span style="mso-spacerun:yes">&nbsp; </span>Had Peterson et al. looked at females (notably features like unicolored vs. bicolored grey underparts, blue vs. green shoulders, etc.) they might well have reached a different conclusion.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:14.0pt;mso-bidi-font-size:12.0pt; font-family:Arial;mso-bidi-font-family:"Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-family:Arial;mso-bidi-font-family:"Times New Roman"'>Comments from Remsen</span></u><span style='font-family:Arial;mso-bidi-font-family:"Times New Roman"'>:<span style="mso-spacerun:yes">&nbsp; </span> YES.<span style="mso-spacerun:yes">&nbsp; </span>I am convinced by <span class=SpellE>Donegan s</span> proposal and by Gary s comments. <o:p></o:p></span></p> <p class=MsoNormal><span style='font-family:Arial;mso-bidi-font-family:"Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-family:Arial;mso-bidi-font-family:"Times New Roman"'>Comments from Pacheco</span></u><span style='font-family:Arial;mso-bidi-font-family: "Times New Roman"'>:  YES.<span style="mso-spacerun:yes">&nbsp; </span></span><span class=SpellE><span style='mso-bidi-font-size:14.0pt;font-family:Arial; mso-bidi-font-family:"Times New Roman"'>Pelas</span></span><span style='mso-bidi-font-size:14.0pt;font-family:Arial;mso-bidi-font-family:"Times New Roman"'> <span class=SpellE>mesmas</span> <span class=SpellE>razıes</span> <span class=SpellE>acima</span>. <o:p></o:p></span></p> <p class=MsoNormal><span style='mso-bidi-font-size:14.0pt;font-family:Arial; mso-bidi-font-family:"Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='mso-bidi-font-size:19.0pt;font-family:Arial; mso-bidi-font-family:ArialMT;mso-bidi-language:EN-US;mso-bidi-font-weight:bold'>Comments from Nores</span></u><span style='mso-bidi-font-size:19.0pt;font-family:Arial; mso-bidi-font-family:ArialMT;mso-bidi-language:EN-US;mso-bidi-font-weight:bold'>:  </span><span style='font-family:Arial;mso-bidi-font-family:"Times New Roman"'>YES.<span style="mso-spacerun:yes">&nbsp; </span>I find <span class=SpellE>Donegan s</span> proposal very convincing and Gary s comments very useful. <span class=hps>By</span> <span class=hps>coloration</span> <span class=hps>and</span> <span class=hps>especially</span> by <span class=hps>distribution</span> <span class=hps>with</span> <span class=hps>a zone</span> <span class=hps>of</span> <span class=hps>hybridization</span> it <span class=hps>appears evident that</span> they <span class=hps>are subspecies</span>. <o:p></o:p></span></p> <p class=MsoNormal><span style='font-family:Arial;mso-bidi-font-family:"Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-family:Arial;mso-bidi-font-family:"Times New Roman"'>Comments from Robbins</span></u><span style='font-family:Arial;mso-bidi-font-family: "Times New Roman"'>:  YES, based on both Thomas Donegan and Gary <span class=SpellE>Stiles s</span> comments. <o:p></o:p></span></p> <p class=MsoNormal><span style='font-family:Arial;mso-bidi-font-family:"Times New Roman"'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-family:Arial;mso-bidi-font-family:"Times New Roman"'>Comments from Zimmer</span></u><span style='font-family:Arial;mso-bidi-font-family:"Times New Roman"'>:  YES, </span><span style='font-family:Arial'>for reasons stated by both Thomas and Gary. <o:p></o:p></span></p> <p class=MsoNormal><span style='font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-family:Arial'>Comments from PÈrez-Em·n</span></u><span style='font-family:Arial'>:  YES. Data presented by both Thomas Donegan and Gary <span class=GramE>show</span> that there is no basis to consider these two taxa as different species. It would be interesting to study the potential hybridization zone at the northern end of the Cordillera Central of Colombia, a pattern that might be found to be similar in other codistributed taxa. </span></p> <p class=MsoNormal><span style='font-family:Arial;mso-bidi-font-family:"Times New Roman"'><o:p>&nbsp;</o:p></span></p> </div> </body> </html>