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text-indent:-.25in; font-family:Wingdings;} @list l14:level7 {mso-level-number-format:bullet; mso-level-text:\F0B7; mso-level-tab-stop:3.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Symbol;} @list l14:level8 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:4.0in; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l14:level9 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:4.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l15 {mso-list-id:180899004; mso-list-type:hybrid; mso-list-template-ids:854778412 1007950612 67698691 67698693 67698689 67698691 67698693 67698689 67698691 67698693;} @list l15:level1 {mso-level-number-format:bullet; mso-level-style-link:"PA Bullet indent 1\.5"; mso-level-text:\F0B7; mso-level-tab-stop:2.0in; mso-level-number-position:left; margin-left:2.0in; text-indent:-.5in; font-family:Symbol;} @list l15:level2 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:1.0in; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l15:level3 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:1.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l15:level4 {mso-level-number-format:bullet; mso-level-text:\F0B7; mso-level-tab-stop:2.0in; mso-level-number-position:left; text-indent:-.25in; font-family:Symbol;} @list l15:level5 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:2.5in; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l15:level6 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:3.0in; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l15:level7 {mso-level-number-format:bullet; mso-level-text:\F0B7; mso-level-tab-stop:3.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Symbol;} @list l15:level8 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:4.0in; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l15:level9 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:4.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l16 {mso-list-id:324600062; mso-list-type:hybrid; mso-list-template-ids:1996631068 1253626790 67698713 67698715 67698703 67698713 67698715 67698703 67698713 67698715;} @list l16:level1 {mso-level-style-link:"PA Num List indent 0\.5"; mso-level-tab-stop:1.0in; mso-level-number-position:left; margin-left:1.0in; text-indent:-.5in;} @list l16:level2 {mso-level-number-format:alpha-lower; mso-level-tab-stop:1.0in; mso-level-number-position:left; text-indent:-.25in;} @list l16:level3 {mso-level-number-format:roman-lower; mso-level-tab-stop:1.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l16:level5 {mso-level-number-format:alpha-lower; mso-level-tab-stop:2.5in; mso-level-number-position:left; text-indent:-.25in;} @list l16:level6 {mso-level-number-format:roman-lower; mso-level-tab-stop:3.0in; mso-level-number-position:right; text-indent:-9.0pt;} @list l16:level8 {mso-level-number-format:alpha-lower; mso-level-tab-stop:4.0in; mso-level-number-position:left; text-indent:-.25in;} @list l16:level9 {mso-level-number-format:roman-lower; mso-level-tab-stop:4.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l17 {mso-list-id:460073160; mso-list-type:hybrid; mso-list-template-ids:1549424892 -1541111700 67698691 67698693 67698689 67698691 67698693 67698689 67698691 67698693;} @list l17:level1 {mso-level-number-format:bullet; mso-level-style-link:"PA Dash List"; mso-level-text:-; mso-level-tab-stop:.5in; mso-level-number-position:left; text-indent:-.5in; font-family:"Times New Roman";} @list l17:level2 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:1.0in; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l17:level3 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:1.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l17:level4 {mso-level-number-format:bullet; mso-level-text:\F0B7; mso-level-tab-stop:2.0in; mso-level-number-position:left; text-indent:-.25in; font-family:Symbol;} @list l17:level5 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:2.5in; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l17:level6 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:3.0in; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l17:level7 {mso-level-number-format:bullet; mso-level-text:\F0B7; mso-level-tab-stop:3.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Symbol;} @list l17:level8 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:4.0in; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l17:level9 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:4.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l18 {mso-list-id:529955104; mso-list-type:hybrid; mso-list-template-ids:1271144654 1212863746 67698691 67698693 67698689 67698691 67698693 67698689 67698691 67698693;} @list l18:level1 {mso-level-number-format:bullet; mso-level-style-link:"PA Bullet indent 0\.5"; mso-level-text:\F0B7; mso-level-tab-stop:1.0in; mso-level-number-position:left; margin-left:1.0in; text-indent:-.5in; font-family:Symbol;} @list l18:level2 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:1.0in; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l18:level3 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:1.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l18:level4 {mso-level-number-format:bullet; mso-level-text:\F0B7; mso-level-tab-stop:2.0in; mso-level-number-position:left; text-indent:-.25in; font-family:Symbol;} @list l18:level5 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:2.5in; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l18:level6 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:3.0in; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l18:level7 {mso-level-number-format:bullet; mso-level-text:\F0B7; mso-level-tab-stop:3.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Symbol;} @list l18:level8 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:4.0in; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l18:level9 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:4.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l19 {mso-list-id:636183296; mso-list-type:hybrid; mso-list-template-ids:1649411412 2063468064 67698713 67698715 67698703 67698713 67698715 67698703 67698713 67698715;} @list l19:level1 {mso-level-number-format:roman-lower; mso-level-style-link:"PA Roman List indent 2"; mso-level-text:"\(%1\)"; mso-level-tab-stop:2.5in; mso-level-number-position:left; margin-left:2.5in; text-indent:-.5in;} @list l19:level2 {mso-level-number-format:alpha-lower; mso-level-tab-stop:1.0in; mso-level-number-position:left; text-indent:-.25in;} @list l19:level3 {mso-level-number-format:roman-lower; mso-level-tab-stop:1.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l19:level5 {mso-level-number-format:alpha-lower; mso-level-tab-stop:2.5in; mso-level-number-position:left; text-indent:-.25in;} @list l19:level6 {mso-level-number-format:roman-lower; mso-level-tab-stop:3.0in; mso-level-number-position:right; text-indent:-9.0pt;} @list l19:level8 {mso-level-number-format:alpha-lower; mso-level-tab-stop:4.0in; mso-level-number-position:left; text-indent:-.25in;} @list l19:level9 {mso-level-number-format:roman-lower; mso-level-tab-stop:4.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l20 {mso-list-id:638732761; mso-list-type:hybrid; mso-list-template-ids:1433716198 67698711 67698713 67698715 67698703 67698713 67698715 67698703 67698713 67698715;} @list l20:level1 {mso-level-number-format:alpha-lower; mso-level-text:"%1\)"; mso-level-tab-stop:.5in; mso-level-number-position:left; text-indent:-.25in;} @list l20:level2 {mso-level-number-format:alpha-lower; mso-level-tab-stop:1.0in; mso-level-number-position:left; text-indent:-.25in;} @list l20:level3 {mso-level-number-format:roman-lower; mso-level-tab-stop:1.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l20:level5 {mso-level-number-format:alpha-lower; mso-level-tab-stop:2.5in; mso-level-number-position:left; text-indent:-.25in;} @list l20:level6 {mso-level-number-format:roman-lower; mso-level-tab-stop:3.0in; mso-level-number-position:right; text-indent:-9.0pt;} @list l20:level8 {mso-level-number-format:alpha-lower; mso-level-tab-stop:4.0in; mso-level-number-position:left; text-indent:-.25in;} @list l20:level9 {mso-level-number-format:roman-lower; mso-level-tab-stop:4.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l21 {mso-list-id:726027768; mso-list-type:hybrid; mso-list-template-ids:-672096694 -411302168 67698713 67698715 67698703 67698713 67698715 67698703 67698713 67698715;} @list l21:level1 {mso-level-number-format:alpha-lower; mso-level-style-link:"PA Alpha List"; mso-level-text:"%1\)"; mso-level-tab-stop:.5in; mso-level-number-position:left; text-indent:-.5in;} @list l21:level2 {mso-level-number-format:alpha-lower; mso-level-tab-stop:1.0in; mso-level-number-position:left; text-indent:-.25in;} @list l21:level3 {mso-level-number-format:roman-lower; mso-level-tab-stop:1.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l21:level5 {mso-level-number-format:alpha-lower; mso-level-tab-stop:2.5in; mso-level-number-position:left; text-indent:-.25in;} @list l21:level6 {mso-level-number-format:roman-lower; mso-level-tab-stop:3.0in; mso-level-number-position:right; text-indent:-9.0pt;} @list l21:level8 {mso-level-number-format:alpha-lower; mso-level-tab-stop:4.0in; mso-level-number-position:left; text-indent:-.25in;} @list l21:level9 {mso-level-number-format:roman-lower; mso-level-tab-stop:4.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l22 {mso-list-id:882132056; mso-list-type:hybrid; mso-list-template-ids:-755958910 -1247479488 67698713 67698715 67698703 67698713 67698715 67698703 67698713 67698715;} @list l22:level1 {mso-level-text:"%1\)"; mso-level-tab-stop:none; mso-level-number-position:left; margin-left:.75in; text-indent:-.25in;} @list l22:level2 {mso-level-number-format:alpha-lower; mso-level-tab-stop:none; mso-level-number-position:left; margin-left:1.25in; text-indent:-.25in;} @list l22:level3 {mso-level-number-format:roman-lower; mso-level-tab-stop:none; mso-level-number-position:right; margin-left:1.75in; text-indent:-9.0pt;} @list l22:level4 {mso-level-tab-stop:none; mso-level-number-position:left; margin-left:2.25in; text-indent:-.25in;} @list l22:level5 {mso-level-number-format:alpha-lower; mso-level-tab-stop:none; mso-level-number-position:left; margin-left:2.75in; text-indent:-.25in;} @list l22:level6 {mso-level-number-format:roman-lower; mso-level-tab-stop:none; mso-level-number-position:right; margin-left:3.25in; text-indent:-9.0pt;} @list l22:level7 {mso-level-tab-stop:none; mso-level-number-position:left; margin-left:3.75in; text-indent:-.25in;} @list l22:level8 {mso-level-number-format:alpha-lower; mso-level-tab-stop:none; mso-level-number-position:left; margin-left:4.25in; text-indent:-.25in;} @list l22:level9 {mso-level-number-format:roman-lower; mso-level-tab-stop:none; mso-level-number-position:right; margin-left:4.75in; text-indent:-9.0pt;} @list l23 {mso-list-id:895507370; mso-list-type:hybrid; mso-list-template-ids:-1489307902 -419628070 67698713 67698715 67698703 67698713 67698715 67698703 67698713 67698715;} @list l23:level1 {mso-level-style-link:"PA Num List"; mso-level-tab-stop:.5in; mso-level-number-position:left; text-indent:-.5in;} @list l23:level2 {mso-level-number-format:alpha-lower; mso-level-tab-stop:1.0in; mso-level-number-position:left; text-indent:-.25in;} @list l23:level3 {mso-level-number-format:roman-lower; mso-level-tab-stop:1.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l23:level5 {mso-level-number-format:alpha-lower; mso-level-tab-stop:2.5in; mso-level-number-position:left; text-indent:-.25in;} @list l23:level6 {mso-level-number-format:roman-lower; mso-level-tab-stop:3.0in; mso-level-number-position:right; text-indent:-9.0pt;} @list l23:level8 {mso-level-number-format:alpha-lower; mso-level-tab-stop:4.0in; mso-level-number-position:left; text-indent:-.25in;} @list l23:level9 {mso-level-number-format:roman-lower; mso-level-tab-stop:4.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l24 {mso-list-id:988359725; mso-list-type:hybrid; mso-list-template-ids:-257899320 -953147078 67698691 67698693 67698689 67698691 67698693 67698689 67698691 67698693;} @list l24:level1 {mso-level-number-format:bullet; mso-level-style-link:"TOC 8"; mso-level-text:\F0B7; mso-level-tab-stop:none; mso-level-number-position:left; text-indent:-.5in; font-family:Symbol;} @list l24:level2 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:none; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l24:level3 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:none; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l24:level4 {mso-level-number-format:bullet; mso-level-text:\F0B7; mso-level-tab-stop:none; mso-level-number-position:left; text-indent:-.25in; font-family:Symbol;} @list l24:level5 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:none; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l24:level6 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:none; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l24:level7 {mso-level-number-format:bullet; mso-level-text:\F0B7; mso-level-tab-stop:none; mso-level-number-position:left; text-indent:-.25in; font-family:Symbol;} @list l24:level8 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:none; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l24:level9 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:none; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l25 {mso-list-id:1032076222; mso-list-type:hybrid; mso-list-template-ids:1265669936 670759984 67698713 67698715 67698703 67698713 67698715 67698703 67698713 67698715;} @list l25:level1 {mso-level-number-format:roman-lower; mso-level-style-link:"PA Roman List indent 1\.5"; mso-level-text:"\(%1\)"; mso-level-tab-stop:2.0in; mso-level-number-position:left; margin-left:2.0in; text-indent:-.5in;} @list l25:level2 {mso-level-number-format:alpha-lower; mso-level-tab-stop:1.0in; mso-level-number-position:left; text-indent:-.25in;} @list l25:level3 {mso-level-number-format:roman-lower; mso-level-tab-stop:1.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l25:level5 {mso-level-number-format:alpha-lower; mso-level-tab-stop:2.5in; mso-level-number-position:left; text-indent:-.25in;} @list l25:level6 {mso-level-number-format:roman-lower; mso-level-tab-stop:3.0in; mso-level-number-position:right; text-indent:-9.0pt;} @list l25:level8 {mso-level-number-format:alpha-lower; mso-level-tab-stop:4.0in; mso-level-number-position:left; text-indent:-.25in;} @list l25:level9 {mso-level-number-format:roman-lower; mso-level-tab-stop:4.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l26 {mso-list-id:1045566613; mso-list-type:hybrid; mso-list-template-ids:-2105485400 -784796264 67698713 67698715 67698703 67698713 67698715 67698703 67698713 67698715;} @list l26:level1 {mso-level-number-format:alpha-lower; mso-level-style-link:"PA Alpha List indent 1"; mso-level-text:"%1\)"; mso-level-tab-stop:1.5in; mso-level-number-position:left; margin-left:1.5in; text-indent:-.5in;} @list l26:level2 {mso-level-number-format:alpha-lower; mso-level-tab-stop:1.0in; mso-level-number-position:left; text-indent:-.25in;} @list l26:level3 {mso-level-number-format:roman-lower; mso-level-tab-stop:1.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l26:level5 {mso-level-number-format:alpha-lower; mso-level-tab-stop:2.5in; mso-level-number-position:left; text-indent:-.25in;} @list l26:level6 {mso-level-number-format:roman-lower; mso-level-tab-stop:3.0in; mso-level-number-position:right; text-indent:-9.0pt;} @list l26:level8 {mso-level-number-format:alpha-lower; mso-level-tab-stop:4.0in; mso-level-number-position:left; text-indent:-.25in;} @list l26:level9 {mso-level-number-format:roman-lower; mso-level-tab-stop:4.5in; mso-level-number-position:right; text-indent:-9.0pt;} @list l27 {mso-list-id:1063405931; mso-list-type:hybrid; mso-list-template-ids:531777548 134807567 134807577 134807579 134807567 134807577 134807579 134807567 134807577 134807579;} @list l27:level1 {mso-level-tab-stop:none; mso-level-number-position:left; text-indent:-.25in;} @list l27:level2 {mso-level-number-format:alpha-lower; mso-level-tab-stop:none; mso-level-number-position:left; text-indent:-.25in;} @list l27:level3 {mso-level-number-format:roman-lower; 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mso-level-tab-stop:3.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Symbol;} @list l37:level8 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:4.0in; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier New";} @list l37:level9 {mso-level-number-format:bullet; mso-level-text:\F0A7; mso-level-tab-stop:4.5in; mso-level-number-position:left; text-indent:-.25in; font-family:Wingdings;} @list l38 {mso-list-id:1668243753; mso-list-type:hybrid; mso-list-template-ids:-1374227108 651730326 67698713 67698715 67698703 67698713 67698715 67698703 67698713 67698715;} @list l38:level1 {mso-level-number-format:bullet; mso-level-style-link:"Bulleted List"; mso-level-text:\F0B7; mso-level-tab-stop:.5in; mso-level-number-position:left; text-indent:-.5in; font-family:Symbol;} @list l38:level2 {mso-level-number-format:bullet; mso-level-text:o; mso-level-tab-stop:1.0in; mso-level-number-position:left; text-indent:-.25in; font-family:"Courier 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mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent1OddRow {mso-style-name:"Medium List 2 - Accent 1"; mso-table-condition:odd-row; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:#D3DFEE; mso-tstyle-border-top:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent1NWCell {mso-style-name:"Medium List 2 - Accent 1"; mso-table-condition:nw-cell; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white;} table.MsoTableMediumList2Accent1SWCell {mso-style-name:"Medium List 2 - Accent 1"; mso-table-condition:sw-cell; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-border-top:cell-none;} table.MsoTableMediumGrid2Accent1 {mso-style-name:"Medium Grid 2 - Accent 1"; mso-tstyle-rowband-size:1; mso-tstyle-colband-size:1; mso-style-priority:68; mso-style-unhide:no; border:solid #4F81BD 1.0pt; mso-padding-alt:0in 5.4pt 0in 5.4pt; mso-border-insideh:1.0pt solid #4F81BD; mso-border-insidev:1.0pt solid #4F81BD; mso-tstyle-shading:#D3DFEE; mso-para-margin:0in; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:10.0pt; font-family:"Times New Roman"; mso-fareast-font-family:SimHei; color:black;} table.MsoTableMediumGrid2Accent1FirstRow {mso-style-name:"Medium Grid 2 - Accent 1"; mso-table-condition:first-row; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#EDF2F8; color:black; mso-ansi-font-weight:bold; mso-bidi-font-weight:bold;} table.MsoTableMediumGrid2Accent1LastRow {mso-style-name:"Medium Grid 2 - Accent 1"; mso-table-condition:last-row; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:1.5pt solid black; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; color:black; mso-ansi-font-weight:bold; mso-bidi-font-weight:bold;} table.MsoTableMediumGrid2Accent1FirstCol {mso-style-name:"Medium Grid 2 - Accent 1"; mso-table-condition:first-column; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; color:black; mso-ansi-font-weight:bold; mso-bidi-font-weight:bold;} table.MsoTableMediumGrid2Accent1LastCol {mso-style-name:"Medium Grid 2 - Accent 1"; mso-table-condition:last-column; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#DBE5F1; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; color:black; mso-ansi-font-weight:normal; mso-bidi-font-weight:normal;} table.MsoTableMediumGrid2Accent1OddColumn {mso-style-name:"Medium Grid 2 - Accent 1"; mso-table-condition:odd-column; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#A7BFDE;} table.MsoTableMediumGrid2Accent1OddRow {mso-style-name:"Medium Grid 2 - Accent 1"; mso-table-condition:odd-row; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#A7BFDE; mso-tstyle-border-insideh:.75pt solid #4F81BD; mso-tstyle-border-insidev:.75pt solid #4F81BD;} table.MsoTableMediumGrid2Accent1NWCell {mso-style-name:"Medium Grid 2 - Accent 1"; mso-table-condition:nw-cell; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:white;} table.MsoTableMediumList2Accent2 {mso-style-name:"Medium List 2 - Accent 2"; mso-tstyle-rowband-size:1; mso-tstyle-colband-size:1; mso-style-priority:66; mso-style-unhide:no; border:solid #C0504D 1.0pt; mso-padding-alt:0in 5.4pt 0in 5.4pt; mso-para-margin:0in; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:10.0pt; font-family:"Times New Roman"; mso-fareast-font-family:SimHei; color:black;} table.MsoTableMediumList2Accent2FirstRow {mso-style-name:"Medium List 2 - Accent 2"; mso-table-condition:first-row; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:3.0pt solid #C0504D; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; font-size:12.0pt; mso-ansi-font-size:12.0pt; mso-bidi-font-size:12.0pt;} table.MsoTableMediumList2Accent2LastRow {mso-style-name:"Medium List 2 - Accent 2"; mso-table-condition:last-row; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:1.0pt solid #C0504D; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent2FirstCol {mso-style-name:"Medium List 2 - Accent 2"; mso-table-condition:first-column; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:1.0pt solid #C0504D; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent2LastCol {mso-style-name:"Medium List 2 - Accent 2"; mso-table-condition:last-column; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:1.0pt solid #C0504D; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent2OddColumn {mso-style-name:"Medium List 2 - Accent 2"; mso-table-condition:odd-column; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:#EFD3D2; mso-tstyle-border-left:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent2OddRow {mso-style-name:"Medium List 2 - Accent 2"; mso-table-condition:odd-row; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:#EFD3D2; mso-tstyle-border-top:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent2NWCell {mso-style-name:"Medium List 2 - Accent 2"; mso-table-condition:nw-cell; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white;} table.MsoTableMediumList2Accent2SWCell {mso-style-name:"Medium List 2 - Accent 2"; mso-table-condition:sw-cell; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-border-top:cell-none;} table.MsoTableMediumGrid2Accent2 {mso-style-name:"Medium Grid 2 - Accent 2"; mso-tstyle-rowband-size:1; mso-tstyle-colband-size:1; mso-style-priority:68; mso-style-unhide:no; border:solid #C0504D 1.0pt; mso-padding-alt:0in 5.4pt 0in 5.4pt; mso-border-insideh:1.0pt solid #C0504D; mso-border-insidev:1.0pt solid #C0504D; mso-tstyle-shading:#EFD3D2; mso-para-margin:0in; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:10.0pt; font-family:"Times New Roman"; mso-fareast-font-family:SimHei; color:black;} table.MsoTableMediumGrid2Accent2FirstRow {mso-style-name:"Medium Grid 2 - Accent 2"; mso-table-condition:first-row; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#F8EDED; color:black; mso-ansi-font-weight:bold; mso-bidi-font-weight:bold;} table.MsoTableMediumGrid2Accent2LastRow {mso-style-name:"Medium Grid 2 - Accent 2"; mso-table-condition:last-row; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:1.5pt solid black; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; color:black; mso-ansi-font-weight:bold; mso-bidi-font-weight:bold;} table.MsoTableMediumGrid2Accent2FirstCol {mso-style-name:"Medium Grid 2 - Accent 2"; mso-table-condition:first-column; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; color:black; mso-ansi-font-weight:bold; mso-bidi-font-weight:bold;} table.MsoTableMediumGrid2Accent2LastCol {mso-style-name:"Medium Grid 2 - Accent 2"; mso-table-condition:last-column; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#F2DBDB; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; color:black; mso-ansi-font-weight:normal; mso-bidi-font-weight:normal;} table.MsoTableMediumGrid2Accent2OddColumn {mso-style-name:"Medium Grid 2 - Accent 2"; mso-table-condition:odd-column; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#DFA7A6;} table.MsoTableMediumGrid2Accent2OddRow {mso-style-name:"Medium Grid 2 - Accent 2"; mso-table-condition:odd-row; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#DFA7A6; mso-tstyle-border-insideh:.75pt solid #C0504D; mso-tstyle-border-insidev:.75pt solid #C0504D;} table.MsoTableMediumGrid2Accent2NWCell {mso-style-name:"Medium Grid 2 - Accent 2"; mso-table-condition:nw-cell; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:white;} table.MsoTableMediumList2Accent3 {mso-style-name:"Medium List 2 - Accent 3"; mso-tstyle-rowband-size:1; mso-tstyle-colband-size:1; mso-style-priority:66; mso-style-unhide:no; border:solid #9BBB59 1.0pt; mso-padding-alt:0in 5.4pt 0in 5.4pt; mso-para-margin:0in; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:10.0pt; font-family:"Times New Roman"; mso-fareast-font-family:SimHei; color:black;} table.MsoTableMediumList2Accent3FirstRow {mso-style-name:"Medium List 2 - Accent 3"; mso-table-condition:first-row; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:3.0pt solid #9BBB59; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; font-size:12.0pt; mso-ansi-font-size:12.0pt; mso-bidi-font-size:12.0pt;} table.MsoTableMediumList2Accent3LastRow {mso-style-name:"Medium List 2 - Accent 3"; mso-table-condition:last-row; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:1.0pt solid #9BBB59; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent3FirstCol {mso-style-name:"Medium List 2 - Accent 3"; mso-table-condition:first-column; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:1.0pt solid #9BBB59; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent3LastCol {mso-style-name:"Medium List 2 - Accent 3"; mso-table-condition:last-column; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:1.0pt solid #9BBB59; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent3OddColumn {mso-style-name:"Medium List 2 - Accent 3"; mso-table-condition:odd-column; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:#E6EED5; mso-tstyle-border-left:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent3OddRow {mso-style-name:"Medium List 2 - Accent 3"; mso-table-condition:odd-row; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:#E6EED5; mso-tstyle-border-top:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent3NWCell {mso-style-name:"Medium List 2 - Accent 3"; mso-table-condition:nw-cell; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white;} table.MsoTableMediumList2Accent3SWCell {mso-style-name:"Medium List 2 - Accent 3"; mso-table-condition:sw-cell; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-border-top:cell-none;} table.MsoTableMediumGrid2Accent3 {mso-style-name:"Medium Grid 2 - Accent 3"; mso-tstyle-rowband-size:1; mso-tstyle-colband-size:1; mso-style-priority:68; mso-style-unhide:no; border:solid #9BBB59 1.0pt; mso-padding-alt:0in 5.4pt 0in 5.4pt; mso-border-insideh:1.0pt solid #9BBB59; mso-border-insidev:1.0pt solid #9BBB59; mso-tstyle-shading:#E6EED5; mso-para-margin:0in; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:10.0pt; font-family:"Times New Roman"; mso-fareast-font-family:SimHei; color:black;} table.MsoTableMediumGrid2Accent3FirstRow {mso-style-name:"Medium Grid 2 - Accent 3"; mso-table-condition:first-row; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#F5F8EE; color:black; mso-ansi-font-weight:bold; mso-bidi-font-weight:bold;} table.MsoTableMediumGrid2Accent3LastRow {mso-style-name:"Medium Grid 2 - Accent 3"; mso-table-condition:last-row; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:1.5pt solid black; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; color:black; mso-ansi-font-weight:bold; mso-bidi-font-weight:bold;} table.MsoTableMediumGrid2Accent3FirstCol {mso-style-name:"Medium Grid 2 - Accent 3"; mso-table-condition:first-column; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; color:black; mso-ansi-font-weight:bold; mso-bidi-font-weight:bold;} table.MsoTableMediumGrid2Accent3LastCol {mso-style-name:"Medium Grid 2 - Accent 3"; mso-table-condition:last-column; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#EAF1DD; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; color:black; mso-ansi-font-weight:normal; mso-bidi-font-weight:normal;} table.MsoTableMediumGrid2Accent3OddColumn {mso-style-name:"Medium Grid 2 - Accent 3"; mso-table-condition:odd-column; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#CDDDAC;} table.MsoTableMediumGrid2Accent3OddRow {mso-style-name:"Medium Grid 2 - Accent 3"; mso-table-condition:odd-row; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#CDDDAC; mso-tstyle-border-insideh:.75pt solid #9BBB59; mso-tstyle-border-insidev:.75pt solid #9BBB59;} table.MsoTableMediumGrid2Accent3NWCell {mso-style-name:"Medium Grid 2 - Accent 3"; mso-table-condition:nw-cell; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:white;} table.MsoTableMediumList2Accent4 {mso-style-name:"Medium List 2 - Accent 4"; mso-tstyle-rowband-size:1; mso-tstyle-colband-size:1; mso-style-priority:66; mso-style-unhide:no; border:solid #8064A2 1.0pt; mso-padding-alt:0in 5.4pt 0in 5.4pt; mso-para-margin:0in; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:10.0pt; font-family:"Times New Roman"; mso-fareast-font-family:SimHei; color:black;} table.MsoTableMediumList2Accent4FirstRow {mso-style-name:"Medium List 2 - Accent 4"; mso-table-condition:first-row; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:3.0pt solid #8064A2; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; font-size:12.0pt; mso-ansi-font-size:12.0pt; mso-bidi-font-size:12.0pt;} table.MsoTableMediumList2Accent4LastRow {mso-style-name:"Medium List 2 - Accent 4"; mso-table-condition:last-row; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:1.0pt solid #8064A2; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent4FirstCol {mso-style-name:"Medium List 2 - Accent 4"; mso-table-condition:first-column; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:1.0pt solid #8064A2; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent4LastCol {mso-style-name:"Medium List 2 - Accent 4"; mso-table-condition:last-column; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:1.0pt solid #8064A2; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent4OddColumn {mso-style-name:"Medium List 2 - Accent 4"; mso-table-condition:odd-column; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:#DFD8E8; mso-tstyle-border-left:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent4OddRow {mso-style-name:"Medium List 2 - Accent 4"; mso-table-condition:odd-row; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:#DFD8E8; mso-tstyle-border-top:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none;} table.MsoTableMediumList2Accent4NWCell {mso-style-name:"Medium List 2 - Accent 4"; mso-table-condition:nw-cell; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-shading:white;} table.MsoTableMediumList2Accent4SWCell {mso-style-name:"Medium List 2 - Accent 4"; mso-table-condition:sw-cell; mso-style-priority:66; mso-style-unhide:no; mso-tstyle-border-top:cell-none;} table.MsoTableMediumGrid2Accent4 {mso-style-name:"Medium Grid 2 - Accent 4"; mso-tstyle-rowband-size:1; mso-tstyle-colband-size:1; mso-style-priority:68; mso-style-unhide:no; border:solid #8064A2 1.0pt; mso-padding-alt:0in 5.4pt 0in 5.4pt; mso-border-insideh:1.0pt solid #8064A2; mso-border-insidev:1.0pt solid #8064A2; mso-tstyle-shading:#DFD8E8; mso-para-margin:0in; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:10.0pt; font-family:"Times New Roman"; mso-fareast-font-family:SimHei; color:black;} table.MsoTableMediumGrid2Accent4FirstRow {mso-style-name:"Medium Grid 2 - Accent 4"; mso-table-condition:first-row; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#F2EFF6; color:black; mso-ansi-font-weight:bold; mso-bidi-font-weight:bold;} table.MsoTableMediumGrid2Accent4LastRow {mso-style-name:"Medium Grid 2 - Accent 4"; mso-table-condition:last-row; mso-style-priority:68; 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{mso-style-name:"Medium Grid 2 - Accent 6"; mso-table-condition:last-column; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#FDE9D9; mso-tstyle-border-top:cell-none; mso-tstyle-border-left:cell-none; mso-tstyle-border-bottom:cell-none; mso-tstyle-border-right:cell-none; mso-tstyle-border-insideh:cell-none; mso-tstyle-border-insidev:cell-none; color:black; mso-ansi-font-weight:normal; mso-bidi-font-weight:normal;} table.MsoTableMediumGrid2Accent6OddColumn {mso-style-name:"Medium Grid 2 - Accent 6"; mso-table-condition:odd-column; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#FBCAA2;} table.MsoTableMediumGrid2Accent6OddRow {mso-style-name:"Medium Grid 2 - Accent 6"; mso-table-condition:odd-row; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:#FBCAA2; mso-tstyle-border-insideh:.75pt solid #F79646; mso-tstyle-border-insidev:.75pt solid #F79646;} table.MsoTableMediumGrid2Accent6NWCell {mso-style-name:"Medium Grid 2 - Accent 6"; mso-table-condition:nw-cell; mso-style-priority:68; mso-style-unhide:no; mso-tstyle-shading:white;} </style> <![endif]--><!--[if gte mso 9]><xml> <o:shapedefaults v:ext="edit" spidmax="1027"> <o:colormru v:ext="edit" colors="#fff1e7"/> <o:colormenu v:ext="edit" fillcolor="#fff1e7"/> </o:shapedefaults></xml><![endif]--><!--[if gte mso 9]><xml> <o:shapelayout v:ext="edit"> <o:idmap v:ext="edit" data="1"/> </o:shapelayout></xml><![endif]--> </head> <body bgcolor="#FFF1E7" lang=EN-US link=blue vlink=purple style='tab-interval: .5in'> <div class=WordSection1> <p class=MsoNormal align=center style='text-align:center'><a name="OLE_LINK2"></a><a name="OLE_LINK1"><span style='mso-bookmark:OLE_LINK2'><b style='mso-bidi-font-weight: normal'><span style='font-size:18.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></b></span></a></p> <p class=MsoNormal align=center style='text-align:center'><span style='mso-bookmark:OLE_LINK1'><span style='mso-bookmark:OLE_LINK2'><b style='mso-bidi-font-weight:normal'><span style='font-size:18.0pt;font-family: Arial'>Revised Proposal (589) to </span></b></span></span><a href="http://www.museum.lsu.edu/~Remsen/SACCBaseline.htm"><span style='mso-bookmark:OLE_LINK1'><span style='mso-bookmark:OLE_LINK2'><b style='mso-bidi-font-weight:normal'><span style='font-size:18.0pt;font-family: Arial;color:blue'>South American Classification Committee</span></b></span></span></a><span style='mso-bookmark:OLE_LINK1'><span style='mso-bookmark:OLE_LINK2'><span style='font-size:18.0pt;font-family:Arial'><o:p></o:p></span></span></span></p> <span style='mso-bookmark:OLE_LINK2'></span><span style='mso-bookmark:OLE_LINK1'></span> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><b style='mso-bidi-font-weight:normal'><u><span lang=EN-GB style='font-size:14.0pt; font-family:Arial;mso-ansi-language:EN-GB'><o:p><span style='text-decoration: none'>&nbsp;</span></o:p></span></u></b></p> <p class=MsoNormal align=center style='text-align:center;mso-outline-level: 1'><b style='mso-bidi-font-weight:normal'><span style='font-size:16.0pt; font-family:Arial'>Split <i style='mso-bidi-font-style:normal'>Epinecrophylla haematonota</i> into four species<i style='mso-bidi-font-style:normal'><o:p></o:p></i></span></b></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><b style='mso-bidi-font-weight:normal'><span style='font-size:14.0pt;font-family:Arial'>Effect on SACC</span></b><span style='font-size:14.0pt;font-family:Arial'>: If adopted, three species would be added to the South American checklist by splitting <i style='mso-bidi-font-style: normal'>Epinecrophylla haematonota</i> into four species as recommended by Whitney, Isler, Bravo, Aristiz·bal, Schunck, Silveira, and Piacentini (2013). Following a comment to the original proposal and the ensuing discussion amended herein, this revision is split into two parts for the committee's consideration as follows:<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='margin-left:.25in'><span style='font-size:14.0pt; font-family:Arial'>A.<span style="mso-spacerun:yes">&nbsp; </span><i style='mso-bidi-font-style:normal'>E. amazonica</i> (Ihering, 1904)<i style='mso-bidi-font-style:normal'>,</i> <i style='mso-bidi-font-style:normal'>E. haematonota</i> (Sclater, 1857)<i style='mso-bidi-font-style:normal'>, E. pyrrhonota </i>(Sclater and Salvin, 1873)<i style='mso-bidi-font-style:normal'>, </i>and <o:p></o:p></span></p> <p class=MsoNormal style='margin-left:.25in'><span style='font-size:14.0pt; font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='margin-left:.25in'><span style='font-size:14.0pt; font-family:Arial'>B. the newly described <i style='mso-bidi-font-style:normal'>E. dentei</i>.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><b style='mso-bidi-font-weight:normal'><span style='font-size:14.0pt;font-family:Arial'>Background</span></b><span style='font-size:14.0pt;font-family:Arial'>: The Stipple-throated Antwren is a widespread denizen of western Amazonia, occurring both south and north of the Amazon. During survey work in the Aripuan„-Machado interfluvium, Amazonas, Brazil, in 2000-2003, Bret Whitney noted that the vocalizations of <i style='mso-bidi-font-style:normal'>E. haematonota</i> antwrens were audibly distinct from those on the opposite side (left bank) of the Rio Madeira.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='mso-outline-level:1'><b style='mso-bidi-font-weight: normal'><span style='font-size:14.0pt;font-family:Arial'>Newly Published information<o:p></o:p></span></b></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial'>A. To provide a foundation for evaluating the taxonomic status of the Aripuan„-Machado population, molecular and vocal analysis of currently recognized subspecies of <i style='mso-bidi-font-style:normal'>E.</i> <i style='mso-bidi-font-style:normal'>haematonota</i> was undertaken, Maximum-likelihood and Bayesian analyses of mtDNA showed that the Aripuan„-Machado population<i style='mso-bidi-font-style:normal'> </i>was embedded within a large well-supported clade containing four distinctive lineages comprising taxa currently known as <i style='mso-bidi-font-style:normal'>E. spodionota</i>, <i style='mso-bidi-font-style:normal'>E. h. haematonota</i> (including <i style='mso-bidi-font-style:normal'>E. fjeldsaai</i>), <i style='mso-bidi-font-style:normal'>E. h. pyrrhonota</i>, and a fourth lineage including <i style='mso-bidi-font-style:normal'>E. h. amazonica</i> and the Aripuan„-Machado population. The relationships among the four lineages are not well resolved, and they are ~5&#8211;6 % divergent. <i style='mso-bidi-font-style: normal'>E. spodionota</i> has previously been recognized as a distinct species, and differences in loudsong pace support the acknowledgement of the remaining lineages as distinct species. The slowest paced loudsongs are delivered by <i style='mso-bidi-font-style:normal'>amazonica</i>, the fastest by nominate <i style='mso-bidi-font-style:normal'>haematonota</i> and <i style='mso-bidi-font-style: normal'>pyrrhonota</i>, and the pace of the loudsong of the Aripuan„-Machado population<i style='mso-bidi-font-style:normal'> </i>is intermediate. All pace comparisons were significant statistically except for between <i style='mso-bidi-font-style:normal'>h. haematonota</i> and <i style='mso-bidi-font-style: normal'>pyrrhonota</i>. The latter two taxa, however, are distinguished morphologically, especially as the female plumage of <i style='mso-bidi-font-style: normal'>pyrrhonota</i> is most distinct in the complex. Given vocal and morphological differences, the extent of genetic diversity among the lineages and the inability to resolve their relationships, even though not a direct basis for species consideration, indicates that the evolution of the four clades, <i style='mso-bidi-font-style:normal'>spodionota</i>, <i style='mso-bidi-font-style:normal'>pyrrhonota</i>, <i style='mso-bidi-font-style: normal'>haematonota</i>, and the clade containing <i style='mso-bidi-font-style: normal'>amazonica</i> and the Aripuan„-Machado population has proceeded to the species level.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial'>B. The phylogeny showed that the Aripuan„-Machado population was sister to E<i style='mso-bidi-font-style:normal'>. h. amazonica</i>, its neighbor to the west on the opposite bank of the Rio Madeira. Differences in vocalizations and morphology support its description as a new species. The new species, <i style='mso-bidi-font-style:normal'>E. dentei</i>, differs vocally by the significantly faster pace of its loudsong and by a diagnosable distinction in note shape which produces loudsongs of a subtlety different quality to the human ear. Adult females of <i style='mso-bidi-font-style:normal'>dentei</i> are readily distinguished from those of <i style='mso-bidi-font-style:normal'>amazonica</i> by color of the throat and of the belly. Maximum-likelihood and Bayesian analysis showed that <i style='mso-bidi-font-style:normal'>dentei</i> and <i style='mso-bidi-font-style:normal'>amazonica</i> were ~3% divergent. Phenotypic differences of vocalizations and plumage supported by genetic results meet previously established guidelines for species consideration in thamnophilids in parapatry (Isler et al. 1998).<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><b style='mso-bidi-font-weight:normal'><span style='font-size:14.0pt;font-family:Arial'>Recommendation</span></b><span style='font-size:14.0pt;font-family:Arial'>: I recommend a &quot;YES&quot; vote on accepting all four species, including <i style='mso-bidi-font-style:normal'>Epinecrophylla dentei</i>, in the list, based on morphology, vocalizations, and genetic distinctions. The status of <i style='mso-bidi-font-style:normal'>fjeldsaai</i> is considered in a separate recommendation.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='mso-outline-level:1'><b style='mso-bidi-font-weight: normal'><span style='font-size:14.0pt;font-family:Arial'>Literature cited</span></b><span style='font-size:14.0pt;font-family:Arial'>:<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='margin-left:.5in;text-indent:-.5in'><span style='font-size:14.0pt;font-family:Arial'>Isler, M. L., P. R. Isler, and B. M. Whitney.<span style="mso-spacerun:yes">&nbsp; </span>1998.<span style="mso-spacerun:yes">&nbsp; </span>Use of vocalizations to establish species limits in antbirds (Passeriformes; Thamnophilidae).<span style="mso-spacerun:yes">&nbsp; </span>Auk 115:577&#8211;590.<o:p></o:p></span></p> <p class=MsoNormal style='margin-left:.5in;text-indent:-.5in'><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='margin-left:.5in;text-indent:-.5in'><span style='font-size:14.0pt;font-family:Arial'>Whitney, B. M., M. L. Isler, G. A. Bravo, N. Aristiz·bal, F. Schunck, L. F. Silveira, and V. de Q. Piacentini. 2013. A new species of <i style='mso-bidi-font-style:normal'>Epinecrophylla</i> antwren from the Aripuan„-Machado interfluvium in central Amazonian Brazil with revision of the  stipple-throated antwren complex. <i style='mso-bidi-font-style: normal'>In</i>: del Hoyo, J., Elliot, A., Sargatal, J., &amp; Christie, D.A. (Eds.), <i style='mso-bidi-font-style:normal'>Handbook of the Birds of the World</i>.<i style='mso-bidi-font-style:normal'> Special Volume: New Species and Global Index</i>. Lynx Edicions, Barcelona, Spain, pp. 263&#8211;267.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal align=center style='text-align:center;mso-outline-level: 1'><span style='font-size:14.0pt;font-family:Arial'>Morton Isler, November 2013<o:p></o:p></span></p> <p class=MsoNormal align=center style='text-align:center;mso-outline-level: 1'><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal align=center style='text-align:center;mso-outline-level: 1'><span style='font-size:14.0pt;font-family:Arial'>=============================================<o:p></o:p></span></p> <p class=MsoNormal align=center style='text-align:center'><span style='font-size:11.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal style='mso-outline-level:1'><b style='mso-bidi-font-weight: normal'><span style='font-size:14.0pt;font-family:Arial;mso-fareast-language: ES-AR'>Further comments:<o:p></o:p></span></b></p> <p class=MsoNormal><b style='mso-bidi-font-weight:normal'><span style='font-size:14.0pt;font-family:Arial;mso-fareast-language:ES-AR'><o:p>&nbsp;</o:p></span></b></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><u><span style='font-size:14.0pt;font-family:Arial;color:black'>Comments from Thomas Donegan</span></u><span style='font-size:14.0pt;font-family:Arial;color:black'>: Our team who works on the Colombian checklist is presently considering the various HBW splits and new species, such as this.<span style="mso-spacerun:yes">&nbsp; </span>Proposal 589 refers to splitting <i style='mso-bidi-font-style:normal'>E. haematonota</i> into 4 species, i.e. treating <i style='mso-bidi-font-style:normal'>pyrrhonota</i>, <i style='mso-bidi-font-style:normal'>dentei</i> and <i style='mso-bidi-font-style: normal'>amazonica</i> each as species when they were previously considered subspecies or undescribed.<span style="mso-spacerun:yes">&nbsp; </span>I had some queries with the vocal data supporting this approach which have been discussed offline with Mort Isler and Bret Whitney, and we agreed to include an edited version of this discussion here:<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'>At the outset, I should note great appreciation for the work that Bret Whitney and colleagues put into getting all these Amazonian species &quot;out there&quot; in such an informative volume; and in particular for this interesting paper on ex-<i style='mso-bidi-font-style:normal'>Myrmotherulas</i>.<span style="mso-spacerun:yes">&nbsp; </span>I apologise for what may be perceived as a negative tone of commentary. One of the most depressing things about publication is the (generally only) negative feedback one tends to receive in return.<span style="mso-spacerun:yes">&nbsp; </span>To redress that here: congratulations for such a mammoth and impressive effort and piece of work, which will surely put the taxonomy of Amazonian birds on a surer footing; and in particular for a very interesting review of these antwrens.<span style="mso-spacerun:yes">&nbsp; </span>I would though raise the following issues on this proposal:<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:.5in;margin-bottom:.0001pt'><span style='font-size:14.0pt; font-family:Arial;color:black'>1. The only claimed differences in voice between the four proposed species in this group are in song speed.<span style="mso-spacerun:yes">&nbsp; </span>The populations postulated for species rank (<i style='mso-bidi-font-style:normal'>haematonota</i>, <i style='mso-bidi-font-style:normal'>pyrrhonota</i>, <i style='mso-bidi-font-style: normal'>amazonica</i> and <i style='mso-bidi-font-style:normal'>dentei</i>) are claimed to differ from one another &quot;significantly&quot; in song speed (except <i style='mso-bidi-font-style:normal'>pyrrhonota</i> vs. <i style='mso-bidi-font-style:normal'>haematonota</i>).<span style="mso-spacerun:yes">&nbsp; </span>However, the wording used is confusing:  Isler et al. (1998, 2007) (who published a test of diagnosability) are cited in the introduction to the section, as a test that two species  <i style='mso-bidi-font-style:normal'>differed significantly</i> .<span style="mso-spacerun:yes">&nbsp; </span>Then when discussing song speed, the paper states that the  <i style='mso-bidi-font-style:normal'>average pace</i> of the four populations <i style='mso-bidi-font-style:normal'>differ significantly</i> .<span style="mso-spacerun:yes">&nbsp; </span>This wording suggests usage of a Student's <i style='mso-bidi-font-style:normal'>t</i>-test or similar (tests of statistical significance) rather than diagnosability.<span style="mso-spacerun:yes">&nbsp; </span>The authors cited only means for a single variable, with no standard deviation data or information about upper and lower bounds of this variable presented and no information about other variables, so it is not possible to work out whether diagnosis is likely.<span style="mso-spacerun:yes">&nbsp; </span>Whitney et al. (2013) did not state any other quantitative vocal differences (except near-diagnostic note interval length differences between <i style='mso-bidi-font-style:normal'>amazonica</i> and <i style='mso-bidi-font-style:normal'>dentei</i>). Subjective differences in note shape are also cited but were considered diagnostic only  almost (i.e. not entirely?) perfectly.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:.5in;margin-bottom:.0001pt'><span style='font-size:14.0pt; font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:.5in;margin-bottom:.0001pt'><span style='font-size:14.0pt; font-family:Arial;color:black'>2. If the single variable cited is diagnostic between some populations, this level of differentiation falls below that recommended for diagnosing antbird species &#8211; where <i style='mso-bidi-font-style: normal'>three</i> diagnosable differences (not a <i style='mso-bidi-font-style: normal'>single</i> one) in variables for calls or songs is the usual benchmark for sympatric species (Isler et al. 1998).<span style="mso-spacerun:yes">&nbsp; </span>(Calls were not studied here.)<o:p></o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:.5in;margin-bottom:.0001pt'><span style='font-size:14.0pt; font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:.5in;margin-bottom:.0001pt'><span style='font-size:14.0pt; font-family:Arial;color:black'>3. Any splitting then relies heavily on molecular data.<span style="mso-spacerun:yes">&nbsp; </span>The molecular study concludes impressive (&gt;5%) differentiation between various groups in a polytomy: <i style='mso-bidi-font-style:normal'>spodionota</i>; <i style='mso-bidi-font-style:normal'>amazonica+dentei</i>; <i style='mso-bidi-font-style: normal'>haematonota+fjeldsaai</i>; and <i style='mso-bidi-font-style:normal'>pyrrhonota.<span style="mso-spacerun:yes">&nbsp; </span></i>However,<i style='mso-bidi-font-style: normal'> </i>splitting based on<i style='mso-bidi-font-style:normal'> </i>molecular data does not adopt a minimalistic approach to resolving the polytomy, because <i style='mso-bidi-font-style:normal'>dentei</i> groups with vocally similar <i style='mso-bidi-font-style:normal'>amazonica </i>and is not in any separate part of the polytomy but is proposed here for species rank.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:.5in;margin-bottom:.0001pt'><span style='font-size:14.0pt; font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'>There is then a highly favourable factor towards most of the splits advocated in this paper and proposal, which is almost entirely overlooked.<span style="mso-spacerun:yes">&nbsp; </span>Two members of the polytomy, <i style='mso-bidi-font-style:normal'>spodionota</i> and <i style='mso-bidi-font-style: normal'>pyrrhonota</i>, are fully sympatric in the East Andes of Colombia.<span style="mso-spacerun:yes">&nbsp; </span>Salaman et al. (2002) noted as follows:<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:.5in;margin-bottom:.0001pt'><span style='font-size:11.0pt; font-family:Arial;color:black'>&quot;</span><span style='font-size:11.0pt; font-family:Calibri;mso-ascii-theme-font:major-latin;mso-hansi-theme-font:major-latin; mso-bidi-font-family:Arial;color:black'>Myrmotherula spodionota, FOOTHILL ANTWREN: Ten birds were caught (one collected) in both primary and secondary growth at Alto RÌo Hornoyaco; five were captured and collected at Fragua (1000-1400 m); one female and 3 males were caught and collected at 1000-1460 m on the RÌo Rumiyaco. These are the first specimen records from Colombia; Willis (1988) reported sightings at El ParaÌso, Dpto Huila in Apr 1962. The species is otherwise known from the Andean East Slope of Ecuador and Peru from 600-1300 m (Hilty &amp; Brown 1986). Interestingly, extensive data from Ecuador indicate an elevational segregation of M. spodionota from its close relative, the Stipple-throated Antwren M. haematonota (Krabbe et al. 1999). However, <u>both were captured in the same mist-nets at 1100 m at Alto RÌo Hornoyaco</u> although they were not observed foraging together</span><span style='font-size: 11.0pt;font-family:Arial;color:black'>.&quot;<o:p></o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:.5in;margin-bottom:.0001pt'><span style='font-size:11.0pt; font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'>As part of the team that captured and processed these two species following simultaneous capture in the same mist-net, I can confirm the accuracy of the notes in this paper!<span style="mso-spacerun:yes">&nbsp; </span>Interestingly, some Colombian East slope specimens referred to by Salaman et al. (2002) from the contact zone were among those sampled for mtDNA by Whitney et al. (2013).<span style="mso-spacerun:yes">&nbsp; </span>Consistent with observations at sites of sympatry and a lack of noted intergradation in the above study, the molecular differentiation between these forms was supported.<span style="mso-spacerun:yes">&nbsp; </span>(<i style='mso-bidi-font-style:normal'>E. spodionota</i> is already recognised as a separate species by the SACC.)<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'>Lumping all of members of the polytomy is not a viable proposition, due to <i style='mso-bidi-font-style: normal'>spodionota</i> and <i style='mso-bidi-font-style:normal'>pyrrhonota</i> being sympatric and demonstrably good species.<span style="mso-spacerun:yes">&nbsp; </span>The logical next step to analyse species limits would be to determine vocal differentiation between the two sympatric members of the polytomy, and compare these differences to pairwise differences between other taxa in the polytomy.<span style="mso-spacerun:yes">&nbsp; </span>However, the authors did not do this: no vocal data on <i style='mso-bidi-font-style:normal'>spodionota</i> is included.<span style="mso-spacerun:yes">&nbsp; </span>We therefore do not know if this polytomy shows atypically low vocal variation but in a way consistent with the vocal variation between sympatric related antbirds.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'>Reading the data as its stands, splitting the polytomy does not have basis in the vocal data presented using the Isler et al. (1998) model.<span style="mso-spacerun:yes">&nbsp; </span>Splitting of the polytomy is, however, supported by molecular results, assuming that recognising monophyletic species is virtuous, because lumping sympatric <i style='mso-bidi-font-style:normal'>spodionota</i> with <i style='mso-bidi-font-style:normal'>haematonota</i> is not viable.<span style="mso-spacerun:yes">&nbsp; </span>New taxon <i style='mso-bidi-font-style: normal'>dentei</i> then emerges as a weaker candidate for species rank, as a result of it being grouped with <i style='mso-bidi-font-style:normal'>amazonica </i>in molecular data (and not an independent part of the polytomy), parapatric in its distribution with respect to <i style='mso-bidi-font-style:normal'>amazonica</i> (these replacing one another on different sides of an Amazonian river, a typical subspecies distribution pattern) and not demonstrating the level of vocal variation from <i style='mso-bidi-font-style:normal'>amazonica</i> as is demonstrated by sympatric antbirds.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'>I would therefore like to propose splitting this proposal into Part A (splitting <i style='mso-bidi-font-style: normal'>haematonota</i> into three species: (<i style='mso-bidi-font-style: normal'>haematonota, pyrrhonota</i>, and <i style='mso-bidi-font-style:normal'>amazonica</i>), for which I see no reasonable alternative even based on the limited vocal data in Whitney et al. (2013), due to the molecular data and sympatry between two forms in the polytomy and the similar divergence of all of them.<span style="mso-spacerun:yes">&nbsp; </span>Part B (splitting <i style='mso-bidi-font-style: normal'>dentei</i> from <i style='mso-bidi-font-style:normal'>amazonica</i>) should be considered separately and does not seem at all clear-cut based on the data that are currently available.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial;color:black'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'>Additional literature cited:<o:p></o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:31.5pt;margin-bottom:.0001pt;text-indent:-31.5pt'><span style='font-size:14.0pt;font-family:Arial'>Isler, M. L., P. R. Isler, and B. M. Whitney.<span style="mso-spacerun:yes">&nbsp; </span>1998.<span style="mso-spacerun:yes">&nbsp; </span>Use of vocalizations to establish species limits in antbirds (Passeriformes; Thamnophilidae).<span style="mso-spacerun:yes">&nbsp; </span>Auk 115:577&#8211;590.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:31.5pt;margin-bottom:.0001pt;text-indent:-31.5pt'><span lang=ES-CO style='font-size:14.0pt;font-family:Arial;mso-ansi-language:ES-CO'>Salaman PGW, Stiles FG, BohÛrquez CI, Alvarez M, UmaÒa AM, Donegan TM &amp; Cuervo AM. 2002. </span><span style='font-size:14.0pt;font-family:Arial'>New and noteworthy records from the east slope of the Andes of Colombia. Caldasia 24(1): 157-189. <o:p></o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:31.5pt;margin-bottom:.0001pt;text-indent:-31.5pt'><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><u><span style='font-size:14.0pt;font-family:Arial'>Comments from Bret Whitney</span></u><span style='font-size:14.0pt;font-family:Arial'>:  I just read [the comments above] and it makes a lot of sense to look into each of the arguments raised.<span style="mso-spacerun:yes">&nbsp; </span><i style='mso-bidi-font-style:normal'>E. dentei</i> is not a strongly differentiated species, it's a weakly differentiated species.<span style="mso-spacerun:yes">&nbsp; </span>It shows weak diagnosability morphologically and vocally (note structure being &quot;almost&quot; diagnostic means one sample failed perfect taxon allocation), genetic divergence is not huge, and it possesses some ecological characteristics that appear to be unique, but the fact remains that the population was well-sampled (relative to many other phylogeographic studies) and shows near-perfect consistency in all of the above character sets throughout its range.<span style="mso-spacerun:yes">&nbsp; </span>As explained in the paper, one taxonomic treatment option would be to recognize <i style='mso-bidi-font-style:normal'>haematonota, spodionota, pyrrhonota</i>, and <i style='mso-bidi-font-style:normal'>amazonica</i> as species with <i style='mso-bidi-font-style:normal'>dentei</i> as a subspecies of <i style='mso-bidi-font-style:normal'>amazonica</i>.<span style="mso-spacerun:yes">&nbsp; </span>If the committees decide that's the way to go, so be it, you will get no further explanation from me.<span style="mso-spacerun:yes">&nbsp; </span>That said, I continue to believe that the best course, the one most indicative of speciation in this group of birds, be it of long-standing or relatively incipient, is to call them all separate species.<span style="mso-spacerun:yes">&nbsp; </span>It is not possible to make a truly strong case for either species or subspecies rank in this case.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><u><span style='font-size:14.0pt;font-family:Arial'>Comments from Mort Isler</span></u><span style='font-size:14.0pt;font-family:Arial'>:<span style="mso-spacerun:yes">&nbsp; </span> As the author who was responsible for the vocal analysis recommending that <i style='mso-bidi-font-style:normal'>dentei</i> be considered distinct at the species level, I will respond to Thomas Donegan's comments. Before I do, let me say that I thought Donegan's comments to be well thought out and considered, and I greatly appreciate the time and energy that they involved. It is great to have a community of individuals, like Thomas and SACC members, who carefully examine species-level recommendations.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'>In response to Donegan comment 1, I can confirm that tests of diagnosability were performed on the data. Unfortunately, space limitations of the HBW volume did not allow the inclusion of the description of the methodology employed nor the results of the test as we have done in all of our previous papers. Pace, a ratio, is not normally distributed, and consequently our test (using the t distribution) for likelihood of overlap with larger samples could not be employed. The statistical test employed was a non-parametric bootstrap simulation (see Isler and Whitney 2011, p. 4 for more description).<span style="mso-spacerun:yes">&nbsp; </span>The Difference Between Means (DBM) of <i style='mso-bidi-font-style:normal'>dentei</i> and <i style='mso-bidi-font-style: normal'>amazonica</i> resulting from this test was five standard deviations, which is well above an acceptable level for considering the difference to be diagnostic. Results of other pairwise comparisons of taxa in the group were diagnostic at a similar level except for between <i style='mso-bidi-font-style: normal'>pyrrhonota</i> and <i style='mso-bidi-font-style:normal'>haematonota</i> as noted in the paper.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'>In response to Donegan comments 2 and 3, the following recommendation in Isler et al. 1998 (p. 586) is sometimes overlooked: &quot;However, three diagnosable differences in vocal characters, as defined herein, should be considered a point of reference, not a requirement----for parapatric taxa, fewer than three diagnosable vocal characters might be considered an appropriate threshold for species recognition.&quot; With regard to <i style='mso-bidi-font-style:normal'>dentei</i> and <i style='mso-bidi-font-style:normal'>amazonica</i>, a question can be raised as to whether they are parapatric and gene flow between them is possible or whether the rivers that appear to separate them geographically are a sufficient barrier. The answer to that question is not known, but we do know that these headwaters rivers provide a less formidable obstacle than those downstream; how much of an obstacle is the issue. Other considerations supporting the recommendation for species status include the almost perfect distinction in note shape (see Whitney comments), ecological distinctions, and genetic diversity. Although I agree with the Donegan comment on the last named, I think it appropriate to keep genetic distance in mind when the decision is borderline, as it is with the current example.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'>For all the reasons expressed by Donegan and expanded above, when the paper was being written, I struggled with the recommendation, and (obviously) came down on the side of a species recommendation for <i style='mso-bidi-font-style:normal'>dentei</i>).<span style="mso-spacerun:yes">&nbsp; </span>But Bret Whitney, in his response, put it perfectly: although <i style='mso-bidi-font-style:normal'>dentei</i> is a clearly differentiated population, arguments can be made for either species or subspecies rank; and although we came down on the species side, and whatever the committee decides will be acceptable.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'>On the second issue (Part A in the Donegan statement) regarding <i style='mso-bidi-font-style:normal'>haematonota, pyrrhonota</i>, and <i style='mso-bidi-font-style:normal'>amazonica</i>, I would only add one observation.<span style="mso-spacerun:yes">&nbsp; </span>The geographic pattern provided by loudsong pace is informative. The slower paced loudsongs of <i style='mso-bidi-font-style:normal'>amazonica</i> are juxtaposed in between the fastest paced loudsongs of <i style='mso-bidi-font-style:normal'>haematonota</i> (and pyrrhonota) and the intermediate paced ones of <i style='mso-bidi-font-style: normal'>dentei</i>. This pattern clearly obviates the possibility of west to east clinality, and although it is not directly relevant to the <i style='mso-bidi-font-style:normal'>amazonica/dentei</i> species rank issue, it seems to add support that <i style='mso-bidi-font-style:normal'>haematonota</i> and <i style='mso-bidi-font-style:normal'>amazonica</i> be considered specifically distinct.<o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'>Additional literature cited:<o:p></o:p></span></p> <p class=MsoBlockText style='margin-top:0in;margin-right:0in;margin-bottom: 0in;margin-left:.25in;margin-bottom:.0001pt;text-indent:-.25in'><span style='font-size:14.0pt;font-family:Arial'>Isler, M. L., and B. M. Whitney. 2011. Species limits in antbirds (Thamnophilidae): the Scale-backed Antbird (<i style='mso-bidi-font-style:normal'>Willisornis poecilinotus</i>) complex. Wilson Journal of Ornithology 123: 1&#8211;14. <o:p></o:p></span></p> <p class=MsoBlockText style='margin-bottom:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;font-family:Arial'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:14.0pt;font-family:Arial'>Additional Comment from Thomas Donegan</span></u><span style='font-size:14.0pt;font-family: Arial'>: </span><span style='font-size:14.0pt;font-family:Arial;mso-fareast-font-family: SimSun;mso-bidi-font-style:italic'>As to falling back to lack of gene flow, supported by c 3% mtDNA variation and good sampling, many diagnostic subspecies could be subjected to this sort of argumentation. If subspecies are to be a valid category based on diagnosability but small differences, then one has to draw the line somewhere between those showing smaller and larger differences.<span style="mso-spacerun:yes">&nbsp; </span>I have found Isler et al.'s &quot;point of reference&quot; a good basis for this in suboscine birds.<span style="mso-spacerun:yes">&nbsp; </span><i>E. dentei</i> is borderline in this respect, but falls the wrong side of the standard suggested.<span style="mso-spacerun:yes">&nbsp; </span>I am not particularly trying to push subspecies treatment versus species as cases like this end up being pretty subjective, but as the other proposed splits within the proposal are much more certain, I do feel they should at least better be unbundled in this committee process.<o:p></o:p></span></p> <p class=MsoNormal><span style='font-size:14.0pt;font-family:Arial;mso-fareast-font-family: SimSun;mso-bidi-font-style:italic'><o:p>&nbsp;</o:p></span></p> <p class=MsoNormal><u><span style='font-size:14.0pt;font-family:Arial'>Additional Comment from Bret Whitney</span></u><span style='font-size:14.0pt;font-family: Arial'>: </span><span style='font-size:14.0pt;font-family:Arial;mso-fareast-font-family: SimSun;mso-bidi-font-style:italic'>The yardstick suggested by Isler et al. 1998 continues to be a conservative standard, but as Mort pointed out, we never imagined that it would become a requirement, and we included statements to that effect.<span style="mso-spacerun:yes">&nbsp; </span>To the contrary, we knew full well that some well-differentiated, universally recognized groups with very simple vocalizations (e.g. <i>Grallaricula flavirostris</i> complex) were definitely not going to display three diagnostic vocal characters.<span style="mso-spacerun:yes">&nbsp; </span>In <i>Hypocnemis</i>, we documented fairly extensive syntopy of <i>subflava</i> and <i>peruviana</i> in Brazil and Peru, but there is only one diagnostic character separating the vocal repert