A Classification of the Bird
Species of South America
South American Classification
Committee
American Ornithologists' Union
(Part 10)
Part 10. Oscine Passeriformes, B (Motacillidae to
Emberizidae) (below)
_______________________________________________________
Part 1. Rheiformes to Podicipediformes
Part 2. Columbiformes to Caprimulgiformes
Part 3. Apodiformes
Part 4. Opisthocomiformes to Strigiformes
Part 5. Trogoniformes to Psittaciformes
Part 6. Suboscine Passeriformes, A (Sapayoidae to Formicariidae)
Part 7. Suboscine Passeriformes, B (Furnariidae)
Part 8. Suboscine Passeriformes, C (Tyrannidae to Tityridae)
Part 9. Oscine Passeriformes, A (Vireonidae to Sturnidae)
Part 11. Oscine Passeriformes, C (Cardinalidae to end)
PASSERIFORMES
Suborder PASSERES (OSCINES) (continued)
MOTACILLIDAE (PIPITS
AND WAGTAILS) 1
Motacilla alba White
Wagtail (V) 1a
Anthus cervinus Red-throated Pipit (V) 1b
Anthus lutescens Yellowish Pipit 2, 3
Anthus furcatus Short-billed Pipit
Anthus chacoensis Pampas Pipit 4
Anthus correndera Correndera Pipit 5
Anthus nattereri Ochre-breasted Pipit
Anthus hellmayri Hellmayr's Pipit 5
Anthus bogotensis Paramo Pipit
1. Genetic
data (e.g., Sibley & Ahlquist 1990, Groth 1998, Barker et al. 2002, 2004,
Johannson et al. 2008, Treplin et
al. 2008) indicate that this family belongs within the "nine-primaried
oscine" cluster of families, probably most closely related to Fringillidae
or Emberizidae. Linear sequence of
species reflects Voelker (1999).
1a. Photographed in French
Guiana 26 Oct. 2009 (Ingels et al. 2010).
SACC proposal
passed to add to main list. Unpublished photo and sight record from
Trinidad (ffrench 1991, Kenefick & Hayes 2006, Ingels et al. 2010).
1b. Photographed
in Ecuador in 2008 (Brinkhuizen et al. 2010). SACC proposal passed to add to main list.
2. Anthus
lutescens was formerly (e.g., Zimmer 1953c) know as A. chii, but see
Hellmayr (1934) and Meyer de Schauensee (1966).
3. The
subspecies parvus of Panama was
formerly (e.g., Ridgway 1904) considered a separate species from Anthus lutescens; they were treated as
conspecific by Hellmayr (1935), and this has been followed in all subsequent
classifications.
4. Anthus chacoensis was
described as a subspecies of A. lutescens by Zimmer (1952), but soon
after, additional specimens convinced him (Zimmer 1953c) that it was a separate
species. Straneck (1987) and
Casa–as et al. (2008) found differences between the two in display flights,
vocalizations, and habitat preferences, including elucidating that the species
does not breed in chaco habitat. SACC proposal
passed to change English name to Pampas Pipit.
5. Voelker (1999) found these two
species to be paraphyletic [but action premature?.]
THRAUPIDAE (TANAGERS) 1
Cyanicterus cyanicterus Blue-backed
Tanager
Nemosia pileata Hooded Tanager 3a
Nemosia rourei Cherry-throated Tanager
3a
Compsothraupis loricata Scarlet-throated
Tanager 2
Sericossypha albocristata White-capped
Tanager 3, 3a
Orchesticus abeillei Brown
Tanager
Parkerthraustes humeralis Yellow-shouldered
Grosbeak 162
Catamblyrhynchus diadema Plushcap
58, 58a
Chlorophanes spiza Green Honeycreeper
42, 42a
Iridophanes pulcherrimus Golden-collared
Honeycreeper 43, 43b
Chrysothlypis chrysomelas Black-and-yellow
Tanager 43d, 45
Chrysothlypis salmoni Scarlet-and-white
Tanager 46
Heterospingus xanthopygius Scarlet-browed
Tanager 43c, 43d
Hemithraupis guira Guira Tanager 44,
43d
Hemithraupis ruficapilla Rufous-headed
Tanager 44
Hemithraupis flavicollis Yellow-backed
Tanager
Conirostrum bicolor Bicolored Conebill
48
Conirostrum margaritae Pearly-breasted
Conebill 48
Conirostrum speciosum Chestnut-vented
Conebill 47, 48
Conirostrum leucogenys White-eared
Conebill 48
Conirostrum binghami Giant
Conebill 50
Conirostrum ferrugineiventre White-browed
Conebill 50
Conirostrum sitticolor Blue-backed
Conebill
Conirostrum albifrons Capped
Conebill 48c
Conirostrum tamarugense Tamarugo
Conebill 49
Conirostrum rufum Rufous-browed Conebill
Conirostrum cinereum Cinereous
Conebill 48b
Sicalis citrina Stripe-tailed
Yellow-Finch 93
Sicalis lutea Puna Yellow-Finch
Sicalis uropygialis Bright-rumped
Yellow-Finch 93b
Sicalis luteocephala Citron-headed
Yellow-Finch
Sicalis auriventris Greater Yellow-Finch
Sicalis olivascens Greenish Yellow-Finch
94, 94a
Sicalis mendozae Monte Yellow-Finch
94, 94a
Sicalis lebruni Patagonian Yellow-Finch
94, 94b
Sicalis columbiana Orange-fronted
Yellow-Finch
Sicalis flaveola Saffron Finch 95,
96
Sicalis luteola Grassland Yellow-Finch
97, 98
Sicalis raimondii Raimondi's Yellow-Finch
98
Sicalis taczanowskii Sulphur-throated
Finch 99
Phrygilus atriceps Black-hooded
Sierra-Finch 68, 69, 70, 71
Phrygilus punensis Peruvian Sierra-Finch
69, 70
Phrygilus gayi Gray-hooded Sierra-Finch
69, 70
Phrygilus patagonicus Patagonian
Sierra-Finch 70
Phrygilus fruticeti Mourning Sierra-Finch
71
Phrygilus dorsalis Red-backed Sierra-Finch
72, 74
Phrygilus erythronotus White-throated
Sierra-Finch 72, 74
Phrygilus carbonarius Carbonated
Sierra-Finch 73
Phrygilus alaudinus Band-tailed Sierra-Finch
Phrygilus unicolor Plumbeous
Sierra-Finch 71
Phrygilus plebejus Ash-breasted
Sierra-Finch 71
Porphyrospiza caerulescens Blue
Finch 67
Melanodera melanodera White-bridled
Finch 77, 78
Melanodera xanthogramma Yellow-bridled
Finch
Catamenia analis Band-tailed Seedeater
138, 93
Catamenia inornata Plain-colored Seedeater
Catamenia homochroa Paramo Seedeater
139
Diglossa gloriosissima Chestnut-bellied
Flowerpiercer 54
Diglossa lafresnayii Glossy
Flowerpiercer 54
Diglossa mystacalis Moustached Flowerpiercer
54
Diglossa gloriosa Merida Flowerpiercer
55
Diglossa humeralis Black Flowerpiercer
55
Diglossa brunneiventris Black-throated
Flowerpiercer 55
Diglossa carbonaria Gray-bellied
Flowerpiercer 55
Diglossa venezuelensis Venezuelan
Flowerpiercer 56
Diglossa albilatera White-sided Flowerpiercer
56
Diglossa duidae Scaled Flowerpiercer
Diglossa major Greater Flowerpiercer
Diglossa indigotica Indigo Flowerpiercer
57
Diglossa sittoides Rusty Flowerpiercer
52, 52a, 53
Diglossa glauca Deep-blue Flowerpiercer
57, 57a
Diglossa caerulescens Bluish
Flowerpiercer 55, 57
Diglossa cyanea Masked Flowerpiercer
55, 57, 57b
Idiopsar brachyurus Short-tailed Finch
74, 75
Xenodacnis parina Tit-like Dacnis
51, 51a
Haplospiza rustica Slaty Finch 79,
80
Haplospiza unicolor Uniform Finch 79,
80
Volatinia jacarina Blue-black Grassquit
106, 107, 93
Conothraupis speculigera Black-and-white
Tanager 1c
Conothraupis mesoleuca Cone-billed
Tanager 1c, 1cc
Creurgops verticalis Rufous-crested
Tanager 4
Creurgops dentatus Slaty Tanager 4,
4a, 4b, 4c
Tachyphonus cristatus Flame-crested
Tanager 14, 14a
Tachyphonus rufiventer Yellow-crested
Tanager 14c
Tachyphonus surinamus Fulvous-crested
Tanager 14
Tachyphonus luctuosus White-shouldered
Tanager 14
Tachyphonus delatrii Tawny-crested
Tanager
Tachyphonus coronatus Ruby-crowned
Tanager 14, 14d
Tachyphonus rufus White-lined Tanager
14d
Tachyphonus phoenicius Red-shouldered
Tanager 14d
Eucometis penicillata Gray-headed
Tanager 14
Trichothraupis melanops Black-goggled
Tanager 14
Coryphospingus pileatus Pileated
Finch 141, 142, 143
Coryphospingus cucullatus Red-crested
Finch 142, 144
Ramphocelus nigrogularis Masked
Crimson Tanager 14
Ramphocelus dimidiatus Crimson-backed
Tanager 16
Ramphocelus melanogaster Black-bellied
Tanager 16, 16a
Ramphocelus carbo Silver-beaked Tanager
16, 16b
Ramphocelus bresilius Brazilian
Tanager 16, 16c
Ramphocelus flammigerus Flame-rumped
Tanager 17, 17a
Lanio fulvus Fulvous Shrike-Tanager
14, 15
Lanio versicolor White-winged
Shrike-Tanager 15
Rhodospingus cruentus Crimson-breasted
Finch 145
Charitospiza eucosma Coal-crested
Finch 107, 140
Cyanerpes nitidus Short-billed
Honeycreeper 39
Cyanerpes lucidus Shining Honeycreeper
41a
Cyanerpes caeruleus Purple Honeycreeper
41a
Cyanerpes cyaneus Red-legged Honeycreeper
Tersina viridis Swallow Tanager
38
Dacnis albiventris White-bellied Dacnis
39
Dacnis lineata Black-faced Dacnis 40
Dacnis flaviventer Yellow-bellied Dacnis
Dacnis hartlaubi Turquoise Dacnis
41
Dacnis nigripes Black-legged Dacnis
Dacnis venusta Scarlet-thighed Dacnis
Dacnis cayana Blue Dacnis
Dacnis viguieri Viridian Dacnis
Dacnis berlepschi Scarlet-breasted Dacnis
Sporophila bouvronides Lesson's
Seedeater 115
Sporophila lineola Lined Seedeater
115
Sporophila leucoptera White-bellied
Seedeater 119
Sporophila peruviana Parrot-billed
Seedeater 120
Sporophila telasco Chestnut-throated
Seedeater 122
Sporophila simplex Drab Seedeater
Sporophila castaneiventris Chestnut-bellied
Seedeater 118
Sporophila minuta Ruddy-breasted Seedeater
118, 122, 123
Sporophila nigrorufa Black-and-tawny
Seedeater
Sporophila bouvreuil Copper
Seedeater 118, 121
Sporophila pileata Pearly-bellied Seedeater
118, 121
Sporophila hypoxantha Tawny-bellied
Seedeater 118, 123
Sporophila ruficollis Dark-throated
Seedeater 118, 124, 125
Sporophila palustris Marsh Seedeater
118, 125
Sporophila hypochroma Rufous-rumped
Seedeater 118, 126
Sporophila cinnamomea Chestnut
Seedeater 118, 126, 127
Sporophila melanogaster Black-bellied
Seedeater 118
Sporophila funerea Thick-billed Seed-Finch
107, 109, 128, 129, 93
Sporophila angolensis Chestnut-bellied
Seed-Finch 129
Sporophila maximiliani Great-billed
Seed-Finch 130, 132
Sporophila crassirostris Large-billed
Seed-Finch 130, 131
Sporophila atrirostris Black-billed
Seed-Finch 130
Sporophila corvina Variable Seedeater
113, 114
Sporophila intermedia Gray
Seedeater 112
Sporophila americana Wing-barred
Seedeater 113
Sporophila murallae Caqueta Seedeater 113
Sporophila fringilloides White-naped
Seedeater 107, 37b, 93
Sporophila luctuosa Black-and-white
Seedeater
Sporophila nigricollis Yellow-bellied
Seedeater 116
Sporophila ardesiaca Dubois's
Seedeater 117
Sporophila caerulescens Double-collared
Seedeater
Sporophila schistacea Slate-colored
Seedeater 111
Sporophila falcirostris Temminck's
Seedeater 111
Sporophila frontalis Buffy-fronted
Seedeater 107, 108, 109, 110, 93
Sporophila plumbea Plumbeous Seedeater
111a
Sporophila beltoni Tropeiro Seedeater
111a
Sporophila collaris Rusty-collared Seedeater
Sporophila albogularis White-throated
Seedeater
Saltatricula
multicolor
Many-colored Chaco Finch 164, 172
Saltatricula
atricollis
Black-throated Saltator 164, 171, 172
Saltator
maximus
Buff-throated Saltator
Saltator
atripennis
Black-winged Saltator
Saltator
orenocensis
Orinocan Saltator
Saltator
coerulescens
Grayish Saltator 166, 167
Saltator
striatipectus
Streaked Saltator 170
Saltator
similis
Green-winged Saltator 166
Saltator
nigriceps
Black-cowled Saltator 169
Saltator
maxillosus
Thick-billed Saltator 166, 168
Saltator
aurantiirostris
Golden-billed Saltator 169
Saltator
cinctus
Masked Saltator 171
Saltator
grossus
Slate-colored Grosbeak 163, 165
Saltator
fuliginosus
Black-throated Grosbeak 163
Coryphaspiza melanotis Black-masked
Finch
Embernagra platensis Great
Pampa-Finch 102, 103, 104, 93
Embernagra longicauda Pale-throated
Pampa-Finch 103, 105
Emberizoides herbicola Wedge-tailed
Grass-Finch 100, 102, 93
Emberizoides duidae Duida Grass-Finch
100
Emberizoides ypiranganus Lesser
Grass-Finch 101
Xenospingus concolor Slender-billed
Finch
Piezorina cinerea Cinereous Finch
93, 93a
Pseudospingus verticalis Black-headed
Hemispingus 5, 12
Pseudospingus xanthophthalmus Drab
Hemispingus 5, 12
Cnemoscopus rubrirostris Gray-hooded
Bush Tanager 12b, 12c
Castanozoster thoracicus Bay-chested
Warbling-Finch 84
Poospiza goeringi Slaty-backed Hemispingus
5, 84
Poospiza rufosuperciliaris Rufous-browed
Hemispingus 5, 11, 84
Poospiza boliviana Bolivian Warbling-Finch
84
Poospiza hypochondria Rufous-sided
Warbling-Finch 84, 84a, 88
Poospiza ornata Cinnamon Warbling-Finch
84, 85
Poospiza nigrorufa Black-and-rufous
Warbling-Finch 84, 86
Poospiza rubecula Rufous-breasted
Warbling-Finch 84, 85, 92
Poospiza caesar Chestnut-breasted
Mountain-Finch 88, 92
Poospiza hispaniolensis Collared
Warbling-Finch 84, 88
Poospiza garleppi Cochabamba
Mountain-Finch 84, 91, 92
Poospiza baeri Tucuman Mountain-Finch
84, 91, 92
Kleinothraupis reyi Gray-capped Hemispingus
Kleinothraupis atropileus Black-capped
Hemispingus 5, 6, 6a
Kleinothraupis parodii Parodi's
Hemispingus 5, 7
Kleinothraupis calophrys Orange-browed
Hemispingus 5, 6,
Sphenopsis frontalis Oleaginous
Hemispingus 5, 9
Sphenopsis melanotis Black-eared
Hemispingus 5, 10
Thlypopsis sordida Orange-headed Tanager
12e
Thlypopsis inornata Buff-bellied Tanager
12e
Thlypopsis fulviceps Fulvous-headed
Tanager
Thlypopsis pyrrhocoma Chestnut-headed
Tanager 12f
Thlypopsis ruficeps Rust-and-yellow Tanager
Thlypopsis superciliaris Superciliaried
Hemispingus 5, 8, 8a, 8b
Thlypopsis ornata Rufous-chested Tanager
12d
Thlypopsis pectoralis Brown-flanked
Tanager 12d
Microspingus alticola Plain-tailed
Warbling-Finch 12a, 84
Microspingus erythrophrys Rusty-browed
Warbling-Finch 12a, 84, 85
Microspingus lateralis Buff-throated
Warbling-Finch 12a, 84, 87
Microspingus cabanisi Gray-throated
Warbling-Finch 12a, 84, 87
Microspingus torquatus Ringed
Warbling-Finch 12a, 84, 88, 89
Microspingus melanoleucus Black-capped
Warbling-Finch 12a, 84, 90
Microspingus cinereus Cinereous
Warbling-Finch 12a, 84, 90
Microspingus trifasciatus Three-striped
Hemispingus 12a
Nephelornis oneilli Pardusco 13
Urothraupis stolzmanni Black-backed
Bush Tanager 59
Cypsnagra hirundinacea White-rumped
Tanager
Donacospiza albifrons Long-tailed
Reed Finch
Incaspiza pulchra Great Inca-Finch
83
Incaspiza personata Rufous-backed Inca-Finch
83
Incaspiza ortizi Gray-winged Inca-Finch
83
Incaspiza laeta Buff-bridled Inca-Finch
Incaspiza watkinsi Little Inca-Finch
Coereba flaveola Bananaquit 147
Tiaris olivaceus Yellow-faced Grassquit
148, 149
Tiaris obscurus Dull-colored Grassquit
150, 22
Tiaris fuliginosus Sooty Grassquit
150
Tiaris bicolor Black-faced Grassquit
151
Certhidea olivacea Green Warbler-Finch 152, 153, 154, 155
Certhidea fusca Gray Warbler-Finch 154
Platyspiza crassirostris Vegetarian
Finch 152
Camarhynchus pallidus Woodpecker
Finch 156
Camarhynchus psittacula Large
Tree-Finch 157
Camarhynchus pauper Medium Tree-Finch
Camarhynchus parvulus Small
Tree-Finch 155
Camarhynchus heliobates Mangrove
Finch 156
Geospiza fuliginosa Small Ground-Finch
Geospiza magnirostris Large
Ground-Finch
Geospiza difficilis Sharp-beaked
Ground-Finch 158, 159
Geospiza scandens Common Cactus-Finch
Geospiza fortis Medium Ground-Finch
160
Geospiza conirostris Large
Cactus-Finch 161
Chlorochrysa phoenicotis Glistening-green
Tanager 27a, 28
Chlorochrysa calliparaea Orange-eared
Tanager 28
Chlorochrysa nitidissima Multicolored
Tanager
Lophospingus pusillus Black-crested
Finch 81, 82, 58c
Lophospingus griseocristatus Gray-crested
Finch
Neothraupis fasciata White-banded
Tanager 1a
Diuca diuca Common Diuca-Finch 76
Diuca speculifera White-winged Diuca-Finch
75, 76
Gubernatrix cristata Yellow
Cardinal 146
Paroaria coronata Red-crested Cardinal
64, 65, 66b
Paroaria dominicana Red-cowled Cardinal
65, 65a
Paroaria nigrogenis Masked Cardinal
66
Paroaria gularis Red-capped Cardinal
66
Paroaria baeri Crimson-fronted Cardinal
66
Paroaria capitata Yellow-billed Cardinal
66
Stephanophorus diadematus Diademed
Tanager
Schistochlamys melanopis Black-faced
Tanager 1a
Schistochlamys ruficapillus Cinnamon
Tanager
Cissopis leverianus Magpie Tanager
1a, 1b
Calochaetes coccineus Vermilion
Tanager 24b
Iridosornis porphyrocephalus Purplish-mantled
Tanager 26, 26b
Iridosornis analis Yellow-throated Tanager
26
Iridosornis jelskii Golden-collared Tanager
Iridosornis rufivertex Golden-crowned
Tanager 27
Iridosornis reinhardti Yellow-scarfed
Tanager 27
Pipraeidea melanonota Fawn-breasted
Tanager 17b
Pipraeidea bonariensis Blue-and-yellow
Tanager 19b
Pseudosaltator rufiventris Rufous-bellied
Mountain-Tanager 24c, 25b
Dubusia taeniata Buff-breasted
Mountain-Tanager 24b, 24c, 25
Dubusia castaneoventris Chestnut-bellied
Mountain-Tanager 24b, 25a
Anisognathus melanogenys Black-cheeked
Mountain-Tanager 23, 23a, 23b, 24b, 24c
Anisognathus lacrymosus Lacrimose
Mountain-Tanager 23a, 23b, 23c
Anisognathus igniventris Scarlet-bellied
Mountain-Tanager 23b
Anisognathus somptuosus Blue-winged
Mountain-Tanager 24, 24a
Anisognathus notabilis Black-chinned
Mountain-Tanager 24a
Buthraupis montana Hooded Mountain-Tanager 24b,
24c
Buthraupis wetmorei Masked Mountain-Tanager
22a
Chlorornis riefferii Grass-green
Tanager 24b
Cnemathraupis eximia Black-chested
Mountain-Tanager 21a, 24c
Cnemathraupis aureodorsalis Golden-backed
Mountain-Tanager 21a, 22
Wetmorethraupis sterrhopteron Orange-throated
Tanager 21
Bangsia arcaei Blue-and-gold Tanager
20a
Bangsia melanochlamys Black-and-gold
Tanager 20
Bangsia rothschildi Golden-chested Tanager
Bangsia edwardsi Moss-backed Tanager
Bangsia aureocincta Gold-ringed Tanager
Tangara ruficervix Golden-naped Tanager
30b, 30c
Tangara cyanoptera Black-headed Tanager
36, 36a, 36aa
Tangara viridicollis Silvery
Tanager 33, 34
Tangara heinei Black-capped Tanager 33
Tangara argyrofenges Green-throated
Tanager 33, 35
Tangara phillipsi Sira Tanager
32, 33
Tangara palmeri Gray-and-gold Tanager 28ab
Tangara peruviana Black-backed Tanager
29, 30a
Tangara preciosa Chestnut-backed Tanager
29, 29a, 30a, 30aa
Tangara meyerdeschauenseei Green-capped
Tanager 30, 30a
Tangara cayana Burnished-buff Tanager
30a, 30aa
Tangara vitriolina Scrub Tanager 29b,
30a
Tangara nigrocincta Masked Tanager 31
Tangara larvata Golden-hooded Tanager
31, 31a
Tangara cyanicollis Blue-necked Tanager
Tangara vassorii Blue-and-black Tanager
32a
Tangara nigroviridis Beryl-spangled
Tanager 32a
Tangara labradorides Metallic-green
Tanager 30c
Tangara cyanotis Blue-browed Tanager
30c
Tangara inornata Plain-colored Tanager
28a, 28aa, 28aa
Tangara mexicana Turquoise Tanager
28aa, 28f
Tangara chilensis Paradise Tanager
28bb, 37a
Tangara velia Opal-rumped Tanager 37,
37a
Tangara callophrys Opal-crowned Tanager
37a
Tangara seledon Green-headed Tanager
28b, 28bb
Tangara fastuosa Seven-colored Tanager
28b, 28bb
Tangara cyanocephala Red-necked
Tanager 28b
Tangara desmaresti Brassy-breasted Tanager
28b, 28bbbb
Tangara cyanoventris Gilt-edged
Tanager 28b, 28bbbb
Tangara lavinia Rufous-winged Tanager
28d
Tangara gyrola Bay-headed Tanager 28d
Tangara rufigenis Rufous-cheeked Tanager
28d
Tangara chrysotis Golden-eared Tanager
28bbb
Tangara xanthocephala Saffron-crowned
Tanager 28bbb
Tangara parzudakii Flame-faced Tanager
28bbb
Tangara schrankii Green-and-gold Tanager
28bbb
Tangara johannae Blue-whiskered Tanager
28bbb
Tangara arthus Golden Tanager 28bbb
Tangara florida Emerald Tanager
28bbb
Tangara icterocephala Silver-throated
Tanager 28bbb
Thraupis cyanocephala Blue-capped
Tanager 19a, 19aa
Thraupis episcopus Blue-gray Tanager
17b, 17c, 28a
Thraupis sayaca Sayaca Tanager 18
Thraupis glaucocolpa Glaucous
Tanager 19
Thraupis cyanoptera Azure-shouldered Tanager
Thraupis ornata Golden-chevroned Tanager
Thraupis palmarum Palm Tanager
Ixothraupis varia Dotted Tanager 28cc
Ixothraupis rufigula Rufous-throated
Tanager 28cc
Ixothraupis guttata Speckled Tanager
28cc, 28e
Ixothraupis xanthogastra Yellow-bellied
Tanager 28c, 28cc
Ixothraupis punctata Spotted
Tanager 28cc, 28ccc
Orthogonys chloricterus Olive-green
Tanager 1d
Lamprospiza melanoleuca Red-billed
Pied Tanager 1d
1.
As traditionally constituted (e.g., Storer 1969, Meyer de Schauensee 1970,
Ridgely & Tudor 1989), the family Thraupidae was strikingly polyphyletic
with respect to the Emberizidae, Fringillidae, and Cardinalidae. Genetic data have revolutionized our understanding
of what constitutes a true tanager more so than for any other New World
family. In fact, the English name
ÒtanagerÓ no longer has any taxonomic meaning (currently associated with
species also in the Emberizidae and Cardinalidae) but rather refers to a
morphotype, usually a bill shape, considered intermediate between the thick
bills of emberizid sparrows and cardinaline grosbeaks and the thin bills of
wood-warblers. The genera Euphonia
and Chlorophonia, traditionally placed in this family, have been
transferred to the Fringillidae (see below). The genera Piranga and Habia have
been found to members of the Cardinalidae (see that family), and Parkerthraustes of the Cardinalidae has
been found to be a member of the Thraupidae (see below). Most members of the former family
ÒCoerebidaeÓ are now known to be tanagers.
More than 30 genera formerly placed in the Emberizidae have been found
to be tanagers, and Chlorospingus,
formerly considered a tanager, has been found to be a member of the Emberizidae. With the transfer of a major block of
former Emberizidae to the Thraupidae (see SACC proposal and references
therein), the current linear sequence is temporary because those genera have
simply been inserted at the end of the family until a phylogenetic linear
sequence can be worked out.
<Remsen and Burns working on this>. The linear sequence through Urothraupis has been adapted from Burns
(1997, 1998), Burns (2002), Burns & Naoki (2004), and Klicka et al.
(2007). Many genera formerly placed
in the Emberizidae are now placed in this family on the basis of those papers
and Barker et al. (2013). The
linear sequence in the Thraupidae follows that in Dickinson & Christidis
(2014) pending a formal proposal to revise it.
1a.
Genetic data (Burns et al. 2003, Burns & Naoki 2004) indicate that Schistochlamys,
Cissopis, and Neothraupis are each other's closest relatives.
1b.
Cissopis is masculine, so the correct spelling of the species name is leverianus
(David & Gosselin 2002b).
1c.
Storer (1960)<> suspected that Conothraupis was closely related to
Sporophila based on remarkable plumage similarities; Zimmer (1947) had
previously suspected that Conothraupis was a "finch" and not a
"tanager" based on bill shape.
1cc.
Conothraupis mesoleuca was described in the monotypic genus Rhynchothraupis,
but see Bond (1951a) for placement in Conothraupis, as in Hellmayr (1936) and most subsequent classifications.
1d.
Some genetic data (Yuri & Mindell 2002) fail to find support for inclusion
of Lamprospiza in Thraupidae; its placement in the Thraupidae was
questioned by (REFS). Barker et al.
(2013) found that it formed a monophyletic group with Orthogonys and Mitrospingus
and that these three genera have to be recognized in a separate family,
Mitrospingidae, to keep related families monophyletic; this was followed by
Dickinson & Christidis (2014). SACC proposal needed
2.
Compsothraupis has been included by some (e.g., Zimmer 1947, Meyer de
Schauensee 1966, 1970) in Sericossypha, but Storer (1970a) suspected
that their similarities represented convergence.
3.
Although Sericossypha albocristata was at one time suspected of not
being a tanager (e.g., Meyer de Schauensee 1966), morphological (Morony 1985)
and genetic data (Burns et al. 2002, 2003) support its traditional placement in
the Thraupidae.
3a.
Genetic data (Burns et al. 2003) indicate that Sericossypha and Nemosia
are closely related and probably sister taxa.
4.
The two species of Creurgops form a superspecies (Dickinson &
Christidis 2014).
4a.
Creurgops dentatus was formerly (e.g., Hellmayr 1936) placed in a
separate genus, Malacothraupis.
4b.
Creurgops is masculine, so the correct spelling of the species name is dentatus
(David & Gosselin 2002b).
4c.
"Malacothraupis gustavi," known from southern Peru and
northern Bolivia and treated reluctantly as a valid species by Hellmayr (1936),
is now known to be a synonym (male plumage) of Creurgops dentatus (Bond
& Meyer de Schauensee 1941, Zimmer 1947b), as suspected by Hellmayr (1936).
See Hybrids and Dubious Taxa.
5. Despite concerns over the monophyly of the genus Hemispingus owing to rather disparate morphology, genetic data (Garc’a-Moreno et al. 2001) provided some support for monophyly for the taxa for which genetic samples are available, including the most morphologically divergent species, H. rufosuperciliaris. A subsequent analysis of a larger data set, however, could not confirm or reject monophyly of the genus (Garc’a-Moreno & FjeldsŒ 2003). Some authors (e.g., Ridgway 1902) considered Hemispingus and Cnemoscopus to be members of the Parulidae, but recent genetic data (Burns REF, Klicka et al. 2007) corroborate that they are correctly placed in the Thraupidae. The comprehensive phylogenies of Burns et al. (2014) and Barker et al. (2015) found that Hemispingus is indeed not monophyletic. Therefore, Burns et al. (2016) (1) described a new genus, Kleinothraupis, for H. atropileus, H. calophrys, and H. parodii; (2) recommended resurrecting Sphenopsis Sclater for H. melanotis and H. frontalis (as in Dickinson & Christidis 2014); and (3) transferred H. superciliaris to Thlypopsis. SACC proposals passed to recognize Kleinothraupis and Sphenopsis. SACC proposal needed on English names.
6.
Hemispingus calophrys was formerly (e.g., Hellmayr
1936, Meyer de Schauensee 1970, Storer 1970a) considered a subspecies of
H. atropileus, but Weske & Terborgh (1974) provided rationale
for treating southern calophrys as a species separate from H.
atropileus; this treatment has been followed by most recent authors
(e.g., Ridgely & Tudor 1989, Sibley & Monroe 1990), but not by Isler
& Isler (1987). Inclusion of calophrys
in H. atropileus would make that broad species paraphyletic with respect
to H. parodii, the sister taxon of H. calophrys (Garc’a-Moreno
& FjeldsŒ 2003). The subspecies
auricularis is at least as distinct genetically and morphologically, and
should presumably given equal taxonomic rank (Garc’a-Moreno et al. 2001,
Garc’a-Moreno & FjeldsŒ 2003). SACC proposal
needed. Taxa ranked as
species in this group were considered to form a superspecies by Sibley &
Monroe (1990).
7.
Described since Meyer de Schauensee (1970): Weske & Terborgh (1974).
8.
Genetic data (Garc’a-Moreno et al. 2001, Garc’a-Moreno & FjeldsŒ 2003)
indicate that Hemispingus superciliaris clusters with the group of Hemispingus
that consists of H. verticalis-H. xanthophthalmus; plumage similarities
also suggest that H. reyi belongs in this group. SACC proposal to alter linear sequence did not pass.
8a.
The leucogaster subspecies group of Peru and the subspecies chrysophrys
of Venezuela were formerly (e.g., Hellmayr 1936) considered separate species
from Hemispingus superciliaris, but see Zimmer (1947).
8b.
"Basileuterus zimmeri," described from Venezuela, is now known
to be a synonym of Hemispingus superciliaris chrysophrys (Meyer de
Schauensee 1966). See Hybrids and Dubious Taxa.
9.
Ridgely & Tudor (1989) suspected that the Venezuelan subspecies
collectively (as H. ignobilis) might deserve separate
species status from Hemispingus frontalis; Hilty (2003), however, noted
that their vocalizations and behavior were similar.
9a.
Storer (1970a) suggested that Haplospiza, along with Middle American Acanthidops,
might be more closely related to Diglossa in the Thraupidae than to
other emberizine genera; this is consistent with some genetic data (Burns et
al. 2003), although broader taxon-sampling (Klicka et al. 2007, Mauck &
Burns 2009, Campagna et al. 2011) indicated that Haplospiza, some Phrygilus,
some Sicalis, Diglossa, and Catamenia are phylogenetically
intermingled. In any case, all
genetic data indicate that Haplospiza
is a member of the Thraupidae. SACC proposal passed to move to Thraupidae.
10.
Genetic data (Garc’a-Moreno et al. 2001, Garc’a-Moreno & FjeldsŒ 2003)
indicate that the distinctive taxon piurae, currently treated as a
subspecies of H. melanotis (e.g., Meyer de Schauensee 1970), is more
distant from the latter than is H. frontalis, and that piurae is
basal to frontalis + melanotis; these analyses, however, are based on
only ca. 300 base-pairs of mtDNA. Ridgely & Greenfield (2001) treated piurae
as a separate species from H. melanotis based on plumage and vocal
differences. SACC proposal to recognize piurae as a species
did not pass. Hilty (2011) also
treated piurae as a separate species. Ridgely & Greenfield (2001) and
Hilty (2011) further recognized the subspecies ochraceus as a separate
species based on plumage differences. SACC proposal
needed.
11.
Described since Meyer de Schauensee (1970): Blake & Hocking (1974).
12.
Genetic data (Garc’a-Moreno et al. 2001, Garc’a-Moreno & FjeldsŒ 2003)
support the traditional view based on plumage, morphology, and biogeography
(Parker et al. 1985, FjeldsŒ & Krabbe 1990, Sibley & Monroe 1990) that Hemispingus
verticalis and H. xanthophthalmus are sister species and form a
superspecies; they were formerly (e.g., Hellmayr 1936, Phelps & Phelps
1950a) placed in a separate genus, Pseudospingus, but Zimmer (1947)
merged this into Hemispingus.
However, the comprehensive phylogenies of Burns et al. (2014) and Barker
et al. (2015) provided the basis for the recommendation by Burns et al. (2016)
that Pseudospingus be resurrected for these two species (as in Dickinson
& Christidis 2014). SACC proposal passed to recognize Pseudospingus. SACC proposal needed on English names.
12a. Hemispingus trifasciatus was formerly (e.g., Hellmayr 1936) placed in the monotypic genus Microspingus, but Zimmer (1947) merged this into Hemispingus. Genetic data (Garc’a-Moreno et al. 2001, Garc’a-Moreno & FjeldsŒ 2003) support the continued inclusion of this species in Hemispingus, although its relationships to other taxa within that genus remain uncertain. However, the comprehensive phylogenies of Burns et al. (2014) and Barker et al. (2015) showed that this species is not closely related to Hemispingus. Burns et al. (2016) recommended resurrecting Microspingus and also the transfer of Poospiza cabanisi, P. lateralis, P. erythrophrys, P. alticola, P. torquatus, P. cinerea, and P. melanoleuca to Microspingus (as in Dickinson & Christidis 2014). SACC proposal passed to recognize Microspingus. SACC proposal needed on English names.
12b.
<REF> included Cnemoscopus within Hemispingus, but this has
not been followed by subsequent authors; genetic data (REFS, Burns et al. 2003,
Garc’a-Moreno & FjeldsŒ 2003) are consistent with a close relationship.
12c.
The southern subspecies chrysogaster was formerly (e.g., REF <any
since Berlepsch 1912?>) considered a separate species from Cnemoscopus
rubrirostris.
12d.
Sibley & Monroe (1990) considered Thlypopsis ornata and T.
pectoralis to form a superspecies; they are parapatric sister species that
may be sympatric at some localities (Zimmer 1947b).
12e.
Sibley & Monroe (1990) considered Thlypopsis sordida and T.
inornata to form a superspecies; Meyer de Schauensee (1966) suggested that
they might be best treated as conspecific, but they may be sympatric at some
localities (Zimmer 1947b).
12f.
The comprehensive phylogenies of Burns et al. (2014) and Barker et al. (2015)
showed that Pyrrhocoma is embedded in
Thlypopsis. Burns et al.
(2016) recommended transfer to Thlypopsis, which also forces creation of a new name, Thlypopsis pyrrhocoma,
for this species. SACC proposal passed to merger Pyrrhocoma into Thlypopsis.
13.
Recently described species and genus: Lowery & Tallman (1976). Although
initially uncertain to which family this genus belonged, genetic data indicate
that it is a tanager (Bledsoe 1988, Burns et al. 2002, 2003).
14.
Genetic data (Burns et al. 2003) indicate that Eucometis, Tachyphonus,
and Lanio are closely related and that Coryphospingus, currently
placed in the Emberizidae, is also part of this group. Burns and Racicot (2009), however, found
that Tachyphonus is not a
monophyletic group, with some species (T.
coronatus, T. rufus, and T. phoenicius) more closely related to Ramphocelus than to other Tachyphonus, and T. surinamus more closely related to Eucometis + Trichothraupis than to other Tachyphonus. Burns and
Racicot (2009) confirmed the close relationship of these genera along with Lanio as represented in the traditional
linear sequences of genera within the Thraupidae, including Eucometis and Trichothraupis as sister genera. However, they also found that the
ÒemberizidÓ genera Coryphospingus and
Rhodospingus are members of this
clade, with the latter possibly embedded within one group of Tachyphonus. SACC proposal passed to transfer Rhodospingus and Coryphospingus
to Thraupidae. Burns and Racicot (2009) recommended
broader generic limits in this group to avoid naming as many as three new
genera, but a broadly defined Ramphocelus
and Lanio would produce genera of
with far greater morphological heterogeneity than in traditional passerine
genera. However, the comprehensive
phylogenies of Burns et al. (2014) and Barker et al. (2015) provided the basis
for the recommendation by Burns et al. (2016) that (1) Islerothraupis Burns et al. 2016 be recognized for T. cristatus, T. luctuosus, and T.
rufiventer; (2) Maschalethraupis
Burns et al. 2016 be recognized for T. surinamus; and (3) Chrysocorypha, Burns et al. 2016 be recognized for T.
delatrii. SACC proposals pending.
14a.
"Tachyphonus nattereri," known from two specimens from
southwestern Brazil and treated as a valid species by Hellmayr (1936), Pinto
(1944), and Meyer de Schauensee (1966, 1970), is now usually considered a
subspecies of, or an aberrant individual of, T. cristatus (Zimmer
1945b, Storer 1970a, Ridgely & Tudor 1989). See Hybrids and Dubious Taxa.
14c.
Tachyphonus rufiventer was formerly (e.g., Hellmayr 1936, Pinto 1944) known as Tachyphonus metallactus,
but see Zimmer (1945b) and Meyer de Schauensee (1966).
14d.
Burns and Racicot (2009) found that Tachyphonus
coronatus and T. rufus were sisters, with T.
phoenicius sister to these two (thus consistent with traditional linear
sequences).
15.
Lanio fulvus and L. versicolor form a superspecies (Haffer
1987).
16.
Ramphocelus dimidiatus, R. carbo, R. melanogaster,
and R. bresilius form a superspecies (Novaes 1959, Storer 1970a,
AOU 1983, Sibley & Monroe 1990). Ramphocelus carbo and R.
melanogaster hybridize to an uncertain extent in Peru (Zimmer 1945b),
but they are generally considered separate species because there is no sign of
intergradation between the two. Ramphocelus carbo and R. bresilius
hybridize to an uncertain extent in southeastern Brazil (Meyer de Schauensee
1966). <check Novaes 1959>
Burns and Racicot (2009) noted that levels of sequence divergence
between these pairs are consistent with low levels of gene flow between them
and their continued treatment as species.
They also found the following relationships among cis-Andean Ramphocelus: (dimidiatus + nigrogularis)
+ [bresilius + (carbo + melanogaster)];
the traditional linear sequence is consistent with these relationships.
16a.
Called "Huallaga Tanager" in Ridgely & Tudor (1989) and Hilty
(2011). SACC proposal to change English name did not pass.
16b.
"Ramphocelus ciropalbicaudatus" Frisch, (2007. Nature
Society News, Griggsville 42[5]:13) used this name for a "new
species" of Ramphocelus tanager that he photographed in S‹o Paulo,
Brazil. The description not qualify as a valid description under the rules of
the International Commission of Zoological Nomenclature; furthermore, the
individual bird is most likely a partially leucistic R. carbo. See CBRO
web page: (http://www.cbro.org.br/CBRO/justif.htm#Rhamphocelus%20ciropalbicaudatus%20F). See Hybrids and Dubious Taxa.
16c.
Schodde & Bock (2016) determined that the species epithet is a noun in
apposition and should the name should be Ramphocelus
bresilia. SACC proposal needed.
17.
The taxon icteronotus was formerly (e.g., Hellmayr 1936, Meyer de
Schauensee 1970) considered a separate species from Ramphocelus flammigerus,
but intergradation between them in southwestern Colombia (Chapman 1917, Sibley
1958) led Storer (1970a) to consider them conspecific, and this treatment has
been followed by most authors subsequently (e.g., Ridgely & Tudor 1989,
Sibley & Monroe 1990). However, as noted by Ridgely & Greenfield
(2001), the differences between these two are comparable to those between two Ramphocelus
taxa (passerinii and costaricensis) recently treated as separate
species (Hackett 1996, AOU 1998). [The problem is that the two Middle American
taxa should not have been split acc. to Stiles] SACC proposal
needed.
17a.
Ramphocelus flammigerus forms a superspecies with Middle American
R. passerinii (Meyer de Schauensee 1966, Storer 1970a, AOU 1983,
Sibley & Monroe 1990).
17b. Genetic data (burns et al. 2014)
indicate that Thraupis is embedded
within broadly defined Tangara. To maintain Thraupis, Burns et al. (2016) recommended dividing Tangara
17bb.
Genetic data (Burns et al. 2003, Burns & Naoki 2004) suggest that Thraupis
and Pipraeidea are closely related and presumably sister genera.
However, Thraupis itself is
polyphyletic (Klicka et al. 2007, Sedano & Burns 2010), so a more complete
Thraupidae phylogeny is needed before changes can be made. Sedano & Burns (2010) found that T. bonariensis is the one species of
Thraupis that is sister to Pipraeidea. SACC proposal passed to transfer Thraupis bonariensis into Pipraeidea. This
was followed by Dickinson & Christidis (2014).
17c.
The species name formerly (e.g., REFS, Phelps & Phelps 1950a) for Thraupis
episcopus was virens, but see <REF>.
18.
Zimmer (1944a) suggested that Thraupis sayaca is only a
subspecies of T. episcopus; the population in their area of
contact, boliviana, is somewhat intermediate in plumage; it was
described as a subspecies of T. episcopus but is closer in
plumage to T. sayaca (Gyldenstolpe 1945). However, Gyldenstolpe
(1945) found the two species sympatric in northern Bolivia; also, the two
evidently differ in vocalizations (Ridgely & Tudor 1989). Also, T. glaucocolpa,
roughly as distinct phenotypically from T. episcopus as T.
sayaca is, is broadly sympatric with T. episcopus (Ridgely
& Tudor 1989). Thraupis episcopus, T. sayaca, and T.
glaucocolpa were considered to form a superspecies by Sibley &
Monroe (1990), but T. glaucocolpa cannot be included because of it is
almost completely overlapping in distribution with T. episcopus.
Although Middle American and trans-Andean populations, the cana group, were treated as separate species by Ridgway (19##;
"Blue Tanager"), they have been treated as conspecific since at least
Hellmayr (1936). <track reasons for change> However, the cana
group is superficially similar to T. sayaca, more so than it is to the
pale-shouldered Amazonian episcopus group. Therefore, if sayaca
is treated as a separate species, perhaps the cana group should also be
returned to species rank.
19.
Thraupis glaucocolpa was formerly (e.g., Hellmayr 1936, Meyer de
Schauensee 1970, Storer 1970a) treated as a subspecies of T. sayaca,
but most recent classifications have treated it as a separate species (e.g.,
Meyer de Schauensee & Phelps 1978, Ridgely & Tudor 1989, Sibley &
Monroe 1990).
19a.
Sedano & Burns (2011) found that Thraupis cyanocephala was not a member of Thraupis but was
close to Anisognathus. Burns
et al. (2016) also recommended resurrecting Sporathraupis,
as in Dickinson & Christidis (2014). SACC proposal pending to
resurrect Sporathraupis.
19aa.
The subspecies olivicyanea of the Coastal Range of Venezuela was
formerly (e.g., REF <since Berlepsch 1912?) considered a separate species
from Thraupis cyanocephala, but they evidently intergrade where in
contact (Meyer de Schauensee 1966).
19b.
The Andean subspecies darwinii was formerly (e.g., Chapman 1926, Zimmer
1930) considered a separate species from Thraupis bonariensis.
20.
Storer (1970a) merged Bangsia into Buthraupis, but this has not
been followed by most subsequent authors; see Monroe et al. (1993). They are not sister genera (Sedano &
Burns 2010).
20a. Recently recorded at Cerro Tacarcuna,
Colombia (Ruiz-Ovalle & Hurtado-Guerra (2014).
21.
Storer (1970a) merged Wetmorethraupis into Buthraupis, but this
has not been followed by most subsequent authors; see Isler & Isler
(1987). They are not sister genera
(Sedano & Burns 2010).
21a.
Sibley & Monroe (1990) considered Buthraupis eximia and B.
aureodorsalis to form a superspecies.
Sedano & Burns (2010) confirmed that they are sister species but
also found that they are sister to Chlorornis
riefferii. SACC proposal passed to remove from Buthraupis
and to resurrect the genus Cnemathraupis
for them.
22.
Described since Meyer de Schauensee (1970): Blake & Hocking (1974).
22a.
Buthraupis wetmorei was formerly (e.g., Hellmayr 1936) placed in a
monotypic genus, Tephrophilus.
Sedano & Burns (2010) showed that it was not a member of Buthraupis. Burns et al. (2016) recommended
resurrecting Tephrophilus, as
in Dickinson & Christidis (2014). SACC proposal pending to resurrect Tephrospilus.
23.
Called "Santa Marta Mountain-Tanager" by Ridgely & Tudor (1989). SACC proposal to change English name did not pass.
23a.
Anisognathus melanogenys and A. lacrymosus form a superspecies
(Sibley & Monroe 1990); they were considered conspecific by Hellmayr (1936)
and Storer (1970a). See Donegan and Avenda–o (2010) for evidence supporting
species rank for melanogenys. SACC proposal did not pass to treat melanogenys as a subspecies of lacrymosus.
23b.
Anisognathus melanogenys, A. lacrymosus, and A. igniventris
were formerly (e.g., Hellmayr 1936) placed in the genus Poecilothraupis,
but see Meyer de Schauensee (1966).
23c.
SACC proposal to change spelling of English name to "Lachrymose"
did not pass.
24.
Anisognathus somptuosus was formerly (e.g., Hellmayr 1936, Meyer de
Schauensee 1970, Storer 1970a, Ridgely & Tudor 1989) known as A.
flavinuchus (or flavinucha), but see Sibley & Monroe (1990). Hilty (2011) elevated the subspecies flavinucha to species rank, with the
English name ÒBolivian Mountain-Tanager.Ó
24a.
Anisognathus somptuosus and A. notabilis were formerly (e.g.,
Hellmayr 1936, Zimmer 1944a, Phelps & Phelps 1950a) placed in the genus Compsocoma,
but see Meyer de Schauensee (1966).
Sedano & Burns (2010) found that their inclusion in Anisognathus was
problematic. SACC proposals pending.
24b.
Contrary to traditional linear sequences, genetic data (Burns et al. 2003,
Burns & Naoki 2004, Sedano & Burns 2010) indicate that Chlorornis
is a member of a group of Andean tanager genera that consists of Buthraupis,
Anisognathus, Delothraupis, Dubusia, and Calochaetes
(and presumably also Bangsia and Wetmorethraupis).
24c. The hyphenated English group-name
ÒMountain-TanagerÓ no longer refers to a monophyletic group (e.g. Burns et al.
2016), so the hyphens must be removed.
SACC proposal needed. <<wait on
prop. 730>>
25.
The subspecies carrikeri of the Santa Marta Mountains, was described as
a separate species from Dubusia taeniata, but was treated as a
subspecies of the latter by Meyer de Schauensee (1966) and subsequent
authors. Described as a separate
species and treated as such by Chapman (1926), the southern subspecies stictocephala was treated as a
subspecies of D. taeniata by Hellmayr
(1936) and subsequent authors.
Vocal differences between it and nominate taeniata are pronounced (Robbins et al., unpubl. data). SACC proposal to elevate stictocephala to species rank did not pass.
25a.
Delothraupis was merged into Dubusia by Meyer de Schauensee
(1966) and <REFS>. Genetic data (Burns et al. 2003, Burns & Naoki
2004, Klicka et al. 2007, Sedano & Burns 2010) indicate that Delothraupis
and Dubusia are sister taxa.
SACC proposal passed to merge these two genera.
25b. Hellack and Schnell (1977, REFs)
previously noted that ÒSaltatorÓ
rufiventris was an unusual saltator based on plumage and morphology.
Genetic data (Klicka et al. 2007, Chaves et al. 2013) have revealed that ÒSaltatorÓ rufiventris is definitely
not a saltator but a tanager, closely related to Delothraupis and Dubusia.
SACC proposal passed to move to Thraupidae. Because there is no other genus name available, the species is
maintained here provisionally, as indicated by quotation marks, pending a
proposal to move to Dubusia or naming
of a new genus. Dickinson &
Christidis (2014) transferred it to Dubusia and called it Rufous-bellied
Mountain-Tanager. Burns et al.
(2016) named a new genus for the species: Pseudosaltator. SACC proposal passed to recognize Pseudosaltator and to change English name to Rufous-bellied
Mountain-Tanager.
26.
Iridosornis porphyrocephalus and I. analis form a
superspecies (Sibley & Monroe 1990); evidence for treating them as separate
species is weak (Ridgely & Tudor 1989); they were considered conspecific by
Hellmayr (1936). OrcŽs (1944) reported them to be sympatric in southern
Ecuador, but the locality for I. porphyrocephalus is widely
regarded as dubious (Ridgely & Tudor 1989, Paynter 1993, Ridgely &
Greenfield 2001).
26b.
Iridosornis is masculine, so the correct spelling of the species name is
porphyrocephalus (David & Gosselin 2002b).
27.
Iridosornis reinhardti was considered a subspecies of I. rufivertex
by Hellmayr (1936), Zimmer (1944a), and Storer (1970a); they form a
superspecies (Parker et al. 1985, Sibley & Monroe 1990).
27a.
Although Chlorochrysa is traditionally placed next to Tangara in
linear sequences (e.g., Meyer de Schauensee 1970), genetic data (Burns &
Naoki 2004) indicate that they are not closely related.
28.
Chlorochrysa phoenicotis and C. calliparaea form a superspecies
(REF).
28a.
Despite being one of the largest genera in the world in terms of species
richness, genetic data (Burns and Naoki 2004) strongly support Tangara
as a monophyletic group. The genus Tangara was formerly called Calospiza
(e.g., Hellmayr 1936), and in literature from the 1800s, Calliste; see
Zimmer (1943b) and Meyer de Schauensee (1966). Although Burns and Naoki (2004)
found support for the monophyly of the genus, relationships within the genus
were found to differ from those implied by traditional linear sequences. SACC proposal passed to change linear sequence. Sedano
& Burns (2010), however, found that Tangara
was monophyletic only if Thraupis was
included. SACC
proposal did not pass to modify generic limits. Burns et al. (2014) confirmed this, and Burns et al. (2016) recommended
resurrection of several genera and described two new ones to partition Tangara into monophyletic groups; see
various Notes 28ab, 28cc, and 36.
28aa.
Isler & Isler (1987) proposed that Tangara inornata and T.
mexicana were sister species based on similarities in sociality, feeding
behavior, habitat, and voice. Genetic data (Burns & Naoki 2004) corroborate
this relationship. SACC proposal passed to change linear sequence.
28ab.
Isler & Isler (1987) proposed that Tangara palmeri and Middle
American T. cabanisi were sister species based on similarities in
plumage, habitat, and voice. Burns
et al. (2014) confirmed this with genetic data. Burns et al. (2016) described a new
genus, Poecilostreptus, for these two
species. SACC
proposal pending to recognize Poecilostreptus.
28b.
Storer (1970a) considered Tangara cyanocephala and T. desmaresti
to form a superspecies. Isler & Isler (1987) considered these two to form a
species group with T. cyanoventris. Genetic data (Burns &
Naoki 2004) indicate that Tangara cyanocephala and T. desmaresti
are likely sister species (T. cyanoventris not sampled), and that
they form a monophyletic group with T. seledon and T. fastuosa, which
are also sister species.
28bb.
Isler & Isler (1987) proposed that Tangara chilensis, T. seledon,
and T. fastuosa were sister species based on similarities in plumage,
behavior, habitat, biogeography, and voice. Genetic data (Burns & Naoki
2004), however, do not corroborate this relationship; see Notes 28b and 37b.
28bbb.
Isler & Isler (1987) proposed that Tangara johannae, T. schrankii,
T. florida, T. arthus, T. icterocephala, T.
xanthocephala, T. chrysotis, and T. parzudakii form a species
group based on similarities in plumage and foraging behavior. Genetic data
(Burns & Naoki 2004) confirm this as a monophyletic group.
28bbbb.
"Tangara gouldi," known only from the type specimens from
southeastern Brazil and considered a species by Hellmayr (1936) and Pinto (1944), is considered a hybrid (T.
cyanoventris X T. desmaresti) (Bond 1947, Meyer de Schauensee 1966,
Storer 1970a). See Hybrids and Dubious Taxa.
28c.
Hilty (2003) suspected that the subspecies phelpsi of the Tepuis might
deserve recognition as a separate species from Tangara xanthogastra.
28cc.
Isler & Isler (1987) proposed that Tangara xanthogastra, T. punctata, T.
guttata, T. varia, and T. rufigula form a species group based
on plumage and vocal similarities. Genetic data (Burns & Naoki 2004, Burns
et al. 2014) confirm this as a monophyletic group. Burns et al. (2016)
recommended restoration of Ixothraupis
Bonaparte for these five species. SACC proposal passed to recognize Ixothraupis.
28d.
Storer (1970a) considered Tangara gyrola and T. lavinia to form a
superspecies. Genetic data (Burns & Naoki 2004) confirm this relationship.
Isler & Isler (1987) suggested that T. rufigenis might also be part
of this group based on plumage and habitat similarities.
28dd.
As pointed out by Meyer de Schauensee (1966), the differences among subspecies
groups currently treated included in T. gyrola seem to be at least of
the same magnitude as those between T. gyrola and T. lavinia;
indeed, the viridissima subspecies group and the bangsi
subspecies group were formerly (e.g., Ridgway 1902, Chapman 1925) each treated
as separate species from T. gyrola.
28e.
The species named used for Tangara guttata was formerly (e.g., Hellmayr
1936, Phelps & Phelps 1950a) chrysophrys, but see Meyer de
Schauensee (1966) and Storer (1970a).
28f.
The subspecies brasiliensis was formerly (e.g., Hellmayr 1936) treated
as a separate species from Tangara mexicana, but most classifications
have followed Zimmer (1943c) in treated them as conspecific.
29.
Some authors (e.g., Hellmayr 1936, Sick 1997) have suspected that Tangara
peruviana and T. preciosa are actually color morphs of the same
species, but they are largely parapatric with little or no evidence for
interbreeding (Firme et al. 2007).
29a.
The species named used for Tangara preciosa was formerly (e.g., Hellmayr
1936, Pinto 1944) castanonota, but see
Meyer de Schauensee (1966) and Storer (1970a).
29b.
The species named used for Tangara vitriolina was formerly (e.g.,
Hellmayr 1936) ruficapilla, but see Meyer de Schauensee (1966) and
Storer (1970a).
30.
Described since Meyer de Schauensee (1970): Schulenberg & Binford (1985).
30a.
Isler & Isler (1987) proposed that Tangara cayana, T. peruviana,
T. preciosa, T. meyerdeschauenseei, T. vitriolina, and
West Indian T. cucullata form a species group based on similarities in
plumage and habitat. Genetic data (Burns & Naoki 2004) corroborate the
close relationship among Tangara cayana, T. vitriolina, T.
cucullata, and T. meyerdeschauenseei (T. peruviana and T.
preciosa not sampled).
30aa.
"Tangara arnaulti," known from one aviary specimen and
reluctantly considered a valid species by Hellmayr
(1936) and Meyer de Schauensee (1966); presumably a hybrid (T.
preciosa X T. cayana) (Hellmayr 1936, Bond 1951a, Meyer de
Schauensee 1966, 1970, Storer 1970a, Ingels 1971). See Hybrids and Dubious Taxa.
30b.
The southern subspecies fulvicervix was formerly (e.g., Berlepsch 1912)
considered a separate species from Tangara ruficervix, a treatment that
will almost certainly be restored; they were treated as conspecific by Hellmayr
(1936) and subsequent authors; they differ in more plumage features than do
many pairs of Tangara treated as separate species, yet they show limited
genetic divergence (Burns & Naoki 2004).
30c.
Isler & Isler (1987) proposed that Tangara ruficervix, T.
labradorides, and T. cyanotis formed a species group based on
plumage similarities. Genetic data (Burns & Naoki 2004) do not confirm this
relationship; these three species do not cluster within any of the other
species groups in the genus.
31.
Tangara larvata was formerly (e.g., Hellmayr 1936, Zimmer 1943b,
Meyer de Schauensee 1966, 1970) considered a subspecies of T. nigrocincta,
but most classifications have generally followed Eisenmann (1957) in treating
it as a separate species; see Wetmore et al. (1984) for rationale. Sibley &
Monroe (1990) considered them to form a superspecies. Genetic data (Burns &
Naoki 2004) show that T. larvata is actually more closely related to T.
cyanicollis than either is to T. nigrocincta, thus confirming
treatment of T. nigrocincta as a separate species from T.
larvata and negating treatment of T. larvata and T. nigrocincta
as members of superspecies.
31a.
Formerly known as "Golden-masked Tanager" (e.g., AOU 1983), but see
<Ridgely REF>.
32.
Described since Meyer de Schauensee (1970): Graves & Weske (1987).
32a.
Isler & Isler (1987) proposed that Tangara vassorii and T. nigroviridis
formed a species group with Central American T. dowii and T. fucosa.
Genetic data (Burns & Naoki 2004) confirm their close relationship, with T.
vassorii basal to the other three,
33.
Graves & Weske (1987) proposed that Tangara heinei, T.
phillipsi, T. argyrofenges, and T. viridicollis formed a
monophyletic group. Genetic data (Burns & Naoki 2004) corroborate
the close relationship of Tangara heinei, T. argyrofenges,
and T. viridicollis. Sibley & Monroe (1990) proposed that T. heinei
and T. phillipsi formed a superspecies.
34.
Called "Silver-backed Tanager" by Isler & Isler (1987), Ridgely
& Tudor (1989), Sibley & Monroe (1990), and Ridgely & Greenfield
(2001). SACC proposal to change English name did not pass.
35.
Called "Straw-backed Tanager" by Isler & Isler (1987), Ridgely
& Tudor (1989), Sibley & Monroe (1990), Ridgely & Greenfield (2001),
and Hilty (2011). SACC proposal to change English name did not pass.
36.
Genetic data (Burns & Naoki 2004) indicate that Tangara cyanoptera
is the sister to the T. heinei species complex (see Note 33),
thus corroborating its traditional position in linear sequences. Burns et al. (2014) found that these
two, plus T. larvata, T. nigrocincta, T. cyanicollis, T. preciosa,
T. peruana, T. meyerdeschauenseei, T.
vitriolina, T. cucullatus, T. cayana, T. viridicollis, T. phillipsi,
and T. argyrofenges, formed a
monophyletic group not included in Tangara
s.s.; Burns et al. (2016) described a new genus, Stilpnia, for this group.
SACC proposal pending to recognize Stilpnia.
36a.
Ridgely & Tudor (1989) and Hilty (2003) suspected that the subspecies whitelyi
of the Tepui region might deserve treatment as a separate species from Tangara
cyanoptera.
36aa.
Dickinson & Christidis (2014) noted that cyanoptera was preoccupied in Tangara
by Saltator cyanopterus Vieillot,
1817, and they replaced Tangara cyanoptera with the name Tangara argentea (Lafresnaye, 1843).
SACC proposal needed.
37.
Ridgely & Tudor (1989) suspected that the subspecies cyanomelas of
the Atlantic forest region of Brazil deserves consideration as a separate
species status from Tangara velia.
37a.
Tangara velia and T. callophrys were formerly (e.g., Hellmayr
1936, Zimmer 1943a, Pinto 1944, Phelps &
Phelps 1950a) placed in a separate genus, Tanagrella; similarities in
plumage and bill shape have suggested to most authors (e.g., Isler & Isler
1987) that T. velia and T. callophrys are sister species. Genetic
data (Burns & Naoki 2004) confirm this relationship and also indicate that T.
chilensis is the sister species to these two.
38.
Tersina was formerly placed in its own monotypic family, Tersinidae
(e.g., Hellmayr 1936, Tordoff 1954a, <>George 1962, Meyer de Schauensee
1970), or subfamily or tribe with the tanagers (e.g., Tersininae as in Storer
1970a, or Tersinini as in <REF>); in all these treatments it was
considered the sister taxon to all other tanagers. Genetic data (Bledsoe 1988,
Burns 1997, Burns et al. 2002, 2003) as well as morphological data (Raikow
1978), however, indicate that it is embedded within the tanagers, and most
closely related to Dacnis and Cyanerpes (Burns et al. 2003).
<Sibley & Ahlquist 1973b>
39.
The genera Dacnis and Cyanerpes were formerly (e.g., Meyer de
Schauensee 1970, FjeldsŒ & Krabbe 1990) considered members of a separate
family, the Coerebidae, but they were considered to be tanagers on the basis of
skull structure by Tordoff (1954a) and were placed in the Thraupidae by Storer
(1969, 1970a). This has been followed in most subsequent classifications.
Molecular data indicate that they are sister genera, but embedded within the
tanagers, forming a group with Tersina (Burns et al. 2002, 2003).
39a.
Dacnis albiventris was formerly (e.g., Hellmayr 1935) placed in a
monotypic genus, Hemidacnis, but Zimmer (1942d) merged this into Dacnis.
40. Ridgely & Tudor (1989) pointed out the trans-Andean egregia
group may deserve species rank. Ridgely et al. (2001) considered egregia
a species separate from lineata based on plumage differences and
disjunct range. SACC proposal to recognize Dacnis egregia as a
separate species did not pass because of insufficient published data.
41. Dacnis
hartlaubi was
formerly (e.g., Hellmayr 1936, Meyer
de Schauensee 1966, 1970, Sibley & Monroe 1990) placed in a monotypic
genus, Pseudodacnis and included in the Thraupidae (as "Turquoise
Dacnis-Tanager") when other Dacnis were placed in the Coerebidae.
Storer (1969, 1970a) merged this genus into Dacnis, and subsequent
classifications have followed this treatment.
41a.
Cyanerpes lucidus and C. caeruleus form a
superspecies (AOU 1983, Sibley & Monroe 1990); they were considered
conspecific by Hellmayr (1935), but they are sympatric in northwestern
Colombia.
42.
The genus Chlorophanes was formerly (e.g., Meyer de Schauensee 1970)
considered a member of a separate family, the Coerebidae, but Tordoff (1954a)
considered it a tanager on the basis of skull morphology and Storer (1969,
1970a) placed it in the Thraupidae. Genetic data indicate that it is embedded
within the tanagers; molecular data do not support a close relationship to Dacnis
or Cyanerpes, but its closest relatives remain uncertain (Burns et al.
2002, 2003).
42a.
"Chlorophanes purpurascens," known only from the type specimen
from "Caracas" and considered a valid species by Hellmayr (1935), is
presumably a hybrid. See Hybrids and Dubious Taxa.
43.
Iridophanes pulcherrimus was placed in the genus Tangara by
Storer (1970a) because of its stunning similarity in plumage to T.
cyanoptera, but genetic data (Burns 1997, Burns et al. 2003) indicates that
it may be the sister genus to Chlorophanes, corroborating predictions
from bill shape and behavior (Ridgely & Tudor 1989).
43b.
Iridophanes is masculine, so the correct spelling of the species name is
pulcherrimus (David & Gosselin 2002b).
43c.
Heterospingus xanthopygius forms a superspecies with Central American H.
rubrifrons (AOU 1983, Sibley & Monroe 1990); they were formerly
(e.g., Hellmayr 1936, Storer 1970a) considered conspecific, but most recent
classifications follow Wetmore et al. (1984) in treating them as separate
species because of the radical differences in male plumage; they are not,
however, sympatric (Wetmore et al. 1984; contra Haffer 1975).
43d.
Genetic data (Burns et al. 2003) indicate that Chrysothlypis and Hemithraupis
are closely related and probably sister taxa, and that Heterospingus is
also part of this group.
44.
Hemithraupis guira and H. ruficapilla hybridize to
at least some degree but do not intergrade where they come in contact (Zimmer
1947b, Sick REF?); they are sister species that constitute a superspecies
(Zimmer 1947b, Sibley & Monroe 1990).
45.
Correct spelling for species name is chrysomelas, not chrysomelaena (David & Gosselin 2002a).
45a. [need citation for record for Colombia]
46.
Chrysothlypis salmoni was formerly (e.g., Hellmayr
1936, Meyer de Schauensee 1970) placed in the monotypic genus Erythrothlypis,
but subsequent classifications have followed its merger by Storer (1970a) into Chrysothlypis.
Ridgely & Tudor (1989) pointed out that Chrysothlypis itself could
be merged into Hemithraupis. See Ridgely & Greenfield (2001) for
doubts as to whether Erythrothlypis should be considered congeneric with
Chrysothlypis.
47.
The genus Conirostrum was formerly (e.g., Meyer de Schauensee 1970,
FjeldsŒ & Krabbe 1990) considered a member of a separate family, the
Coerebidae, and by others a member of the Parulidae (Ridgway 1902, Beecher
1951, Tordoff 1954a, Lowery & Monroe 1968). Genetic data (based on C.
speciosum, C. bicolor, and C. sitticolor) indicate that it should be
included in the tanagers, with Oreomanes as the sister genus (Burns et
al. 2002, 2003) or perhaps included within Conirostrum (Lovette &
Bermingham 2002).
48.
Conirostrum speciosum, C. leucogenys, C. bicolor,
and C. margaritae were formerly (e.g., Hellmayr 1935, Tordoff
1954a) placed in a separate genus, Ateleodacnis, but see Zimmer (1942d)
for its merger into Conirostrum. Ridgely & Tudor (1989) pointed out
that this lowland group was quite distinct from montane Conirostrum, perhaps
meriting a return to treatment as a separate genus.
48b.
Ridgely & Greenfield (2001) suggested that the subspecies littorale
of western Peru might deserve recognition as a separate species from Conirostrum
cinereum.
48c.
The atrocyanea subspecies group was formerly (e.g., <REFS, not
Hellmayr 1935>) considered a separate species from Conirostrum albifrons.
49.
Described since Meyer de Schauensee (1970): Johnson & Millie (1972);
described as C. tamarugensis, but see Mayr & Vuilleumier (1983).
50.
The genus Oreomanes was formerly (e.g., Meyer de Schauensee 1970,
FjeldsŒ & Krabbe 1990) considered a member of a separate family, the
Coerebidae, and by others a member of the Parulidae <REF>; it was
tentatively placed in the Thraupidae by Storer (1970a). Wetmore et al. (1984) noted that plumage
similarities suggest a close link between Oreomanes and Poospiza.
Genetic data indicate that it should be included in the tanagers,
with Conirostrum as the sister genus (Burns et al. 2002, 2003). A hybrid
Oreomanes fraseri X C. ferrugineiventre and plumage,
morphological, and foraging similarities also supports their close relationship
(Schulenberg 1985, FjeldsŒ 1992). The
comprehensive phylogenies of Burns et al. (2014) and Barker et al. (2015)
showed that Oreomanes is
embedded within Conirostrum; therefore Burns et al. (2016)
transferred it to Conirostrum; this would also require a change in the
species name (to Conirostrum binghami), as in Dickinson & Christidis (2014). SACC
proposal passed to change to Conirostrum
binghami.
51.
The genus Xenodacnis was formerly (e.g., Meyer de Schauensee 1970,
FjeldsŒ & Krabbe 1990) considered a member of a separate family, the
Coerebidae, but it was placed in the Thraupidae by Storer (1970a); genetic data
indicate that it should be included in the Thraupidae, and that it forms a
group with Acanthidops, Diglossa, and Catamenia (Burns et
al. 2002, 2003); its relationship to other "Coerebidae" was
questioned long ago by Zimmer (1942d). <Beecher>
51a.
The northern petersi subspecies group was formerly (e.g., Bond &
Meyer de Schauensee 1939) considered a separate species from Xenodacnis
parina.
52.
The genus Diglossa was formerly (e.g., Meyer de Schauensee 1970, FjeldsŒ
& Krabbe 1990) considered a member of a separate family, the Coerebidae.
Others considered it a member of the Parulidae (e.g., AOU 1983) or Emberizidae
(REF) because of its apparent close relationship to Acanthidops
(Eisenmann in Meyer de Schauensee 1966), traditionally placed in the
Emberizidae. Tordoff (1954a) considered Diglossa to be a tanager based
on skull morphology, and Storer (1970a) placed it in the Thraupidae; this has
been followed in most subsequent classifications. Genetic data confirm that it
should be included in the tanagers, and forms a group with Acanthidops, Catamenia, and other genera, many formerly considered
emberizine (Burns et al. 2002, 2003, Mauck & Burns 2009). Vuilleumier (1969) divided the genus
into four species groups, but genetic data (Mauck & Burns 2009) indicate
that these are not monophyletic groups.
52a.
The English name "flowerpiercer" is hyphenated in many classifications.
SACC proposal to change to "flower-piercer" did not pass.
53.
Diglossa sittoides was formerly (e.g., Hellmayr 1935, Zimmer 1942e,
Phelps & Phelps 1950a, Meyer de Schauensee 1970, Storer 1970a, Wetmore et
al. 1984, Isler & Isler 1987) considered conspecific with D. baritula
and D. plumbea of Middle America, but see Vuilleumier (1969) and Hackett
(1995) for their treatment as separate species, representing a return to the
classification of <REF ? Ridgway>; they form a superspecies (Sibley &
Monroe 1990), and genetic data (Mauck and Burns 2009) confirm that they form a
monophyletic group: D. sittoides + (D. plumbea + D. baritula).
54.
Diglossa gloriosissima and D. mystacalis were
considered subspecies of D. lafresnayii by many (e.g.,
Hellmayr 1935, Meyer de Schauensee 1970, Storer 1970a, Isler & Isler
1987), but see Vuilleumier (1969) for ranking of these two groups as species
separate from D. lafresnayii.
These three form a superspecies (Vuilleumier 1969, Sibley & Monroe 1990). Genetic data (Mauck and Burns 2009)
confirm that they form a monophyletic group and indicate that D. gloriosissima
and D. lafresnayii are more closely
related than either is to D. mystacalis.
55.
Diglossa gloriosa, D. humeralis, D. brunneiventris,
and D. carbonaria were formerly considered conspecific
("Carbonated Flower-piercer") by many (e.g., Hellmayr 1935, Phelps
& Phelps 1950a, Meyer de Schauensee 1970, Storer 1970a, Isler & Isler
1987), but see Graves (1982) for treatment of all four as separate species.
However, whether the two disjunct populations of D. brunneiventris
are more closely related to each other than to adjacent and intervening D.
humeralis populations remains to be determined. The four species constitute a
superspecies (Vuilleumier 1969, Sibley & Monroe 1990); genetic data (Mauck
and Burns 2009) confirm that they form a monophyletic group.
56.
Diglossa venezuelensis and D. albilatera constitute
a superspecies (Vuilleumier 1969, Isler & Isler 1987, Sibley & Monroe
1990); genetic data (Mauck and Burns 2009) confirm that they form a
monophyletic group.
57.
Diglossa caerulescens was
formerly treated in the monotypic genus Diglossopis, but was included in Diglossa
by Hellmayr (1935). Bock (1985) proposed separating glauca,
caerulescens, and cyanea in the genus Diglossopis, and he
interpreted morphological data to indicate that this and Diglossa were not sister genera. This classification was followed by
Sibley & Monroe (1990), who also added D.
indigotica to the genus based on VuilleumierÕs (1969) species groups. FjeldsŒ & Krabbe (1990), Ridgely
& Greenfield (2001), and (Hilty 2003) also recognized Diglossopis. Genetic data (Mauck & Burns 2009)
indicate that species assigned to Diglossopis
do not form a monophyletic group: D.
cyanea and D. caerulescens are
sister species, and D. glauca is
sister to these two, but D. indigotica
is sister to the D. baritula
superspecies. SACC proposal passed to change linear sequence. SACC proposal to recognize modified Diglossopis did not pass. Genetic
data also indicate that broadly defined Diglossa
is a monophyletic group, contra Bock (1985).
57a.
Called "Golden-eyed Flowerpiercer" in Ridgely & Greenfield
(2001). SACC proposal to change English name did not pass.
57b.
Hilty (2003) noted that differences in song between northern and southern
populations of Diglossa cyanea suggest that two species may be involved.
58.
The genus Catamblyrhynchus is often maintained in a separate monotypic
family, Catamblyrhynchidae (e.g., Hellmayr 1938, Phelps & Phelps 1950a,
Tordoff 1954a, Meyer de Schauensee 1970) or subfamily (Paynter 1970b, Ridgely
& Tudor 1989); genetic data suggest that it should be at least tentatively
included in the Thraupidae (Bledsoe 1988).
58a.
Formerly (e.g., Meyer de Schauensee 1970) called "Plush-capped
Finch".
59.
Genetic data support continued but tentative inclusion of Urothraupis in
the Thraupidae (Bledsoe 1988, Sibley & Ahlquist 1990); some authors have
included it tentatively in the Emberizidae (e.g., Paynter 1970a), based on
superficial similarity to Atlapetes.
64.
The genus Paroaria has been placed traditionally in the Emberizidae,
sometimes with the cardinal grosbeaks (e.g., Hellmayr 1938, Meyer de Schauensee
1966, 1970), which in this classification are considered a separate family,
Cardinalidae. Tordoff (1954a) concluded that it was not a cardinaline but an
emberizine genus, based on skeletal data. Genetic data, however, indicate that
the genus Paroaria belongs in the Thraupidae (Yuri & Mindell 2002,
Burns and Naoki 2004, Klicka et al. 2007), as suspected long ago by Paynter
(1970a). SACC proposal passed to move to Thraupidae.
65.
Paroaria coronata and P. dominicana were considered
to form a superspecies by Sibley & Monroe (1990).
65a.
"Paroaria humberti," described from a captive individual, was
treated as a valid species by Hellmayr (1938), who noted that it could be
simply a melanistic P. dominicana; it has been treated as such by
subsequent authors (e.g., Paynter 1970c). See Hybrids and Dubious Taxa.
66.
Paroaria gularis, P. baeri, and P. capitata
form a superspecies (Sibley & Monroe 1990); evidence for treating them as
separate species is weak (Paynter 1970a); Hellmayr (1938) suspected that P.
capitata might best be treated as a subspecies of P. gularis,
and Meyer de Schauensee (1966) suspected that baeri might also
best be treated as a subspecies of P. gularis. The subspecies P. g. nigrogenis of Venezuela was formerly (e.g., REF) treated
as a separate species from Paroaria gularis. D‡valos & Porzecanski (2009) also
found evidence that nigrogenis was
not most closely to P. gularis; they
elevated all diagnosable taxa to species rank based on the phylogenetic species
concept. SACC
proposal passed to elevate nigrogenis to species rank; SACC proposal to elevate cervicalis and xinguensis
to species rank did not pass. Lopes & Gonzaga (2013) provided
additional rationale for treating xinguensis
as a separate species from P. baeri. SACC proposal to recognize xinguensis as a separate species did not pass.
66b.
<Coccopsis for Paroaria as in Phelps & Phelps 1950a.>
67.
In linear sequences, the genus Porphyrospiza has traditionally (e.g.,
Hellmayr 1938, Meyer de Schauensee 1966, 1970) been associated with the Passerina
buntings, now in the Cardinalidae; Paynter (1970c) even merged Porphyrospiza
into Passerina. This traditional association is based on shared plumage
coloration and pattern with some Passerina buntings. It is generally
agreed, however, among recent authors who know Porphyrospiza caerulescens in
the field (e.g., Ridgely & Tudor 1989, REFS) that it is not related to
cardinaline buntings but to other genera currently in the Emberizidae, as
proposed by Tordoff (1954a) based on skull morphology. Genetic data (Klicka et
al. 2007) confirm that it is not related to cardinalines but rather is the
sister (of the taxa sampled) to Phrygilus alaudinus (but that both are
members of the Thraupidae, not Emberizidae). SACC proposal passed to move to Thraupidae.
68.
Genetic data indicate that all Phrygilus species sampled so far (Burns et al. 2002, 2003, Klicka et al.
2007, Campagna et al. 2011) belong in the Thraupidae. SACC proposal passed to move to Thraupidae.
69.
Phrygilus atriceps is treated as a separate species from P.
gayi because of sympatric breeding reported in Chile (see Johnson 1967).
<check BBOC 109:66-82, 1989> Paynter (1970a) considered the taxon punensis
to be a subspecies of P. atriceps, whereas Meyer de Schauensee
(1970) treated it as a subspecies of P. gayi. Recent
classifications usually follow Ridgely & Tudor (1989), who elevated punensis
to species rank based largely on unpublished data. SACC
proposal to consider punensis and atriceps
conspecific did not pass.
70.
Sibley & Monroe (1990) considered Phrygilus atriceps, P.
punensis, P. gayi, and P. patagonicus to form a
superspecies, but the degree of apparent sympatry between P. gayi
and P. patagonicus would make inclusion of the latter questionable.
71
The genus Phrygilus is highly
polyphyletic (Klicka et al. 2007, Campagna et al. 2011). Campagna et al. (2011) found that the
genus consists of at least four lineages: (1) gayi, patagonicus, punensis, and atriceps, which comprise the sister group to Melanodera (including extralimital Rowettia goughensis); (2) fruticeti,
alaudinus, and carbonaria; (3) plebejus
and unicolor, which are sister to Haplospiza; and (4) dorsalis and erythronotus,
which are sister to Idiopsar. The type species for Phrygilus is gayi. HellmayrÕs (1938)
synonymy indicates that Rhopospina
Cabanis is available for Group 2, with fruticeti
the designated type species, and that Geospizopsis
Bonaparte is available for Group 3, with unicolor
the designated type species.
Clearly, major taxonomic revisions are needed but additional taxon
sampling is needed within the Thraupidae.
SACC proposal to revise classification did not pass. The
comprehensive phylogenies of Burns et al. (2014) and Barker et al. (2015) provided
the basis for the recommendation by Burns et al. (2016) that (1) Rhopospina be resurrected for fruticeti, (2) Corydospiza Sundevall be resurrected for alaudinus and carbonarius,
(3) Geospizopsis Bonaparte
be resurrected for plebejus and unicolor (as in Dickinson
& Christidis 2014), and (4) Ephippiospingus Burns et al. 2016 be recognized for dorsalis and erythronotus. This
would leave gayi, patagonicus, punensis, and atriceps as the only
species in Phrygilus. Dickinson
& Christidis (2014) resurrected Rhopospina
and included in it not only fruticeti,
alaudinus, and carbonarius but also Porphyrospiza caerulescens. SACC proposals pending.
71b.
With the transfer of Phrygilus unicolor to Haplospiza, this species becomes Haplospiza geospizopsis (Bonaparte, 1853) because Haplospiza unicolor Cabanis, 1851, is
the type species for Haplospiza.
72.
Phrygilus dorsalis and P. erythronotus form a
superspecies (Sibley & Monroe 1990). FjeldsŒ & Krabbe (1990) noted
interbreeding between the two and wondered whether dorsalis might not be
just a color phase P. erythronotus.
Genetic data confirm that they are sister species but are not members of
Phrygilus (Burns et al. 2016 and
references therein). Burns et al.
described a new genus, Ephippiospingus, for these
two species. SACC proposals pending.
Dickinson & Christidis
(2014) included them both in Idiopsar.
73.
SACC proposal did not pass to change English name to something other than
"Carbonated" (because
the latter nowadays associated primarily with carbon dioxide injection into
beverages). Dickinson &
Christidis (2014) called it ÒCarbon Finch.Ó
74.
Paynter (1970a) suggested that Idiopsar could be merged into Diuca, but genetic data (Campagna et al. 2011)
indicate that they are distantly related and that Idiopsar is the sister
to Phrygilus erythronotus + P. dorsalis (and thus also a member
of the Thraupidae; see Note 6).
SACC proposal passed to move to Thraupidae.
75.
Genetic data (Bledsoe 1988, Klicka et al. 2007, Campagna et al. 2011) indicate
that Diuca does not belong in the Emberizidae; in fact, it is embedded
within the Òcore tanagersÓ (Sedano & Burns 2010). SACC proposal passed to move to Thraupidae.
76.
Diuca speculifera and D. diuca were considered to form
a superspecies by Paynter (1970a) and Sibley & Monroe (1990), but they are
not closely related (Burns et al. 2014).
Burns et al. (2016) described a new genus, Chionodacryon, for
speculifera. SACC proposal pending to recognize Chionodacryon.
77. Genetic data (Campagna et al. 2011)
indicate that Melanodera is sister to
one of the Phrygilus lineages (see
Note 6), and that it is a member of the Thraupidae. SACC proposal passed to move to Thraupidae.
78.
Called "White-bridled Finch" in Mazar Barnett and Pearman (2001) and
Gill and Wright (2006). The Australian estrildid Poephila cincta is also
known as "Black-throated Finch," the name formerly used by most New
World references. Called
ÒCanary-winged FinchÓ by Jaramillo (2011). SACC proposal passed to change to "Canary-winged
Finch" but subsequent SACC proposal passed to change to "White-bridled Finch."
79. Haplospiza rustica was formerly (e.g., Hellmayr 1938, Phelps & Phelps 1950a) treated in a separate genus, Spodiornis, but Meyer de Schauensee (1966) merged this into Haplospiza. The two species of Haplospiza form a superspecies (Sibley & Monroe 1990); evidence for treating them as separate species was considered weak by Paynter (1970a). However, Burns et al. (2014) found that they are not sister species, with H. rustica sister to Central American Acanthidops bairdi; Burns et al. (2016) recommended resurrection of Spodiornis for H. rustica. Dickinson & Christidis (2014) including all three species in Haplospiza. SACC proposal pending to recognize Spodiornis.
81.
Genetic data (Klicka et al. 2007) indicate that Lophospingus is not a
member of the Emberizidae but belongs in the Thraupidae; in fact, it is
embedded within the Òcore tanagersÓ (Sedano & Burns 2010). SACC proposal passed to move to Thraupidae.
82.
Lophospingus pusillus was formerly placed in the monotypic genus Schistospiza,
but see Miller (1928) for its merger into Lophospingus.
83.
Incaspiza pulchra, I. personata, and I. ortizi
form a superspecies (Sibley & Monroe 1990); I. pulchra and I.
personata were considered conspecific by Hellmayr (1938) and Paynter
(1970a), but recent classifications have followed Meyer de Schauensee (1966) in
treated them as separate species.
84.
Genetic data indicate that Poospiza belongs in the Thraupidae (Lougheed
et al. 2000, Burns et al. 2002, 2003, Klicka et al. 2007, Campagna et al. 2011)
and forms a group with Pyrrhocoma, Thlypopsis, Cypsnagra, Nephelornis,
Hemispingus, and Cnemoscopus. SACC proposal passed to move to Thraupidae. Genetic
data (Lougheed et al. 2000, Klicka et al. 2007) also indicate that Poospiza
is likely polyphyletic. The
comprehensive phylogenies of Burns et al. (2014) and Barker et al. (2015)
confirmed that Poospiza is polyphyletic and should be restricted to P. hispaniolensis, P. rubecula, P. nigrorufa, P. boliviana, and P. ornata. Additionally,
Burns et al. (2016) recommended (1) the transfer of the following species to Poospiza: Hemispingus rufosuperciliaris, H.
goeringi, Compsospiza garleppi and C. baeri; (2) resurrection
of Poospizopsis for P.
caesar and P. hypochondria (see
Note 88); (3) recognition of Castanozoster
Burns et al. 2016 for P. thoracica;
and (4) transfer of remaining species to Microspingus
(see Note 12a). Dickinson &
Christidis (2014) resurrected Orospingus,
Riley, 1922, for rufosuperciliaris
and H. goeringi. SACC proposals passed to revise generic limits. SACC proposal needed on English names.
84a.
Dickinson & Christidis (2014) noted that the original spelling was hypocondria and that hypochondria is an unjustified
emendation. SACC proposal to change to hypocondria
did not pass.
85.
Hellmayr (1938) considered Poospiza erythrophrys and P.
rubecula to be sister species.
FjeldsŒ (1992) proposed that Poospiza erythrophrys and P.
ornata are sister species based on plumage similarities, as reflected in
their placement in traditional linear sequences. Lougheed et al.'s (2000)
limited genetic data did not support such a relationship.
86.
Ridgely & Tudor (1989) and Sibley & Monroe (1990) considered Andean
populations (whitii with wagneri) as a separate species
("Black-and-chestnut Warbling-Finch") from Poospiza nigrorufa,
as suggested by Meyer de Schauensee (1966); Sibley & Monroe (1990) treated
them as species, and members of a superspecies; Mazar Barnett & Pearman
(2001) also treated them as separate species. SACC proposal to recognize whitii as separate
species did not pass.
87.
Poospiza cabanisi has been
traditionally treated as a subspecies of P.
lateralis, although Meyer de Schauensee (1966) and Ridgely & Tudor
(1989) suggested that cabanisi might merit treatment as a separate
species. Assis et al. (2007)
presented evidence for treating cabanisi as a separate species. SACC proposal passed to elevate cabanisi to species rank.
88.
Poospiza caesar was formerly (e.g., Hellmayr
1938, Meyer de Schauensee 1970) placed in a monotypic genus, Poospizopsis;
most authors (e.g., Ridgely & Tudor 1989, Sibley & Monroe 1990) have
followed Paynter (1970a) in merging it into Poospiza, as suggested by
Meyer de Schauensee (1966). Genetic data (Lougheed et al. 2000) indicate that
it is not particularly close to other Poospiza except for P.
hispaniolensis, and that these two are probably sister species. Hellmayr (1938) considered Poospiza
hispaniolensis and P. torquata to be closely related, and Meyer de
Schauensee (1966) suggested that they perhaps should be treated as conspecific;
genetic data, however, indicate that they are only distantly related (Lougheed
et al. 2000). The comprehensive
phylogenies of Burns et al. (2014) and Barker et al. (2015) showed that P. caesar and P. hypochondria are sister species that are not members of Poospiza to Tiaris; Burns et al. (2016) recommended
resurrection of Poospizopsis for these two species (as in Dickinson
& Christidis 2014). SACC proposal pending to recognize Poospizopsis.
89.
Lougheed et al. (2000) suggested that the southern subspecies pectoralis
deserves treatment as a separate species from nominate torquata based on
genetic distance data.
90.
Poospiza melanoleuca has sometimes been treated as a subspecies
of P. cinerea (e.g., Paynter 1970a, FjeldsŒ & Krabbe 1990), but see
Short (1975) and Ridgely & Tudor (1989) for continued treatment as separate
species, as in Hellmayr (1938), Pinto (1944), and
Meyer de Schauensee (1966). Ridgely
& Tudor (1989) noted that melanoleuca has priority over cinerea
if considered conspecific, contra Paynter (1970a).
91.
Compsospiza garleppi and C. baeri were traditionally
(e.g., Hellmayr 1938, Meyer de Schauensee
1966, 1970) placed in that genus, but Paynter (1970a) merged this into Poospiza,
as suggested by Bond (1951), and this treatment has been followed by most
authors subsequently. Genetic data
(Klicka et al. 2007) indicate that baeri is not the sister species to
three other Poospiza sampled.
SACC proposal passed to resurrect Compsospiza.
Compsospiza garleppi
and C. baeri form a superspecies (Sibley & Monroe 1990), and
genetic data (Lougheed et al. 2000) support their treatment as sister species;
Meyer de Schauensee (1966), Paynter (1970a), and FjeldsŒ & Krabbe (1990)
suggested that they might be treated as conspecific. Burns et al. (2016) based on Burns et
al. (2014) recommended that they be transferred back to Poospiza; see Note 84. SACC proposal pending.
92.
FjeldsŒ (1992) proposed that Poospiza caesar, P. rubecula,
P. garleppi, and P. baeri formed a monophyletic
group based on plumage similarities, contrary to their placement in traditional
linear sequences. Lougheed et al.'s (2000) limited genetic data did not support
such a group.
93.
Genetic data indicate that the genus Sicalis (based on S. luteola
or S. olivascens) belongs in the Thraupidae (Bledsoe 1988, Burns et al.
2002, 2003, Klicka et al. 2007). SACC proposal passed to move to Thraupidae. Sicalis
(and Piezorina, Emberizoides, Embernagra, Volatinia, Sporophila,
Oryzoborus [now Sporophila],
Amaurospiza, Dolospingus [now
Sporophila], and Catamenia) was placed with the carduelines by Hellmayr (1938) but then
moved to the emberizines by Meyer de Schauensee (1966) based on the
morphological data of Tordoff (1954). Paynter (1970a) emphasized that species
limits within the genus were uncertain and in need of much study.
93a.
Kashin (1978) noted that the original spelling of the genus by Lafresnaye was Piezorina, and that the subsequent
spelling Piezorhina was an
unjustified emendation, and Gregory & Dickinson (2012) agreed. SACC proposal passed to change to Piezorina.
93b.
Correct spelling is uropigialis
according to Dickinson & Christidis (2014). SACC proposal
needed.
94.
Sibley & Monroe (1990) considered Sicalis olivascens and S.
lebruni to form a superspecies; they were considered conspecific by Paynter
(1970a), as suggested by Meyer de Schauensee (1966).
94a.
Areta et al. (2012) provided evidence for treatment of the subspecies mendozae as a separate species from Sicalis
olivascens. SACC proposal passed to treat mendozae
as separate species.
94b.
L—pez-Lanœs (2017) described a new species, Sicalis
holmbergi, from prov. Buenos Aires, Argentina, from a population previously
considered to be S. lebruni. SACC proposal pending to recognize Sicalis holmbergi.
95.
Called "Saffron Yellow-Finch" in Mazar Barnett & Pearman (2001).
96.
"Sicalis striata," known from two specimens from Prov.
Buenos Aires and treated as a species by (REF), is now considered to be based
on an immature S. flaveola pelzelni (Paynter 1970a, Sibley & Monroe
1990). See Hybrids and Dubious Taxa.
97.
Meyer de Schauensee (1966) and Ridgely & Tudor (1989) suggested that the
southern subspecies luteiventris might represent a separate species from
Sicalis luteola, and it was treated as such by Sibley & Monroe
(1990) and AOU (<?> 1983, 1998). Meyer de Schauensee (1966), Ridgely
& Tudor (1989), and Hilty (2003) also suggested that the subspecies bogotensis
might deserve species rank.
98.
Sibley & Monroe (1990) considered Sicalis luteola and S.
raimondii to form a superspecies; they were formerly (e.g., Hellmayr 1938)
considered conspecific, but Koepcke (1963) found that they were locally
sympatric.
99.
Sicalis taczanowskii was formerly (e.g., Hellmayr
1938, Meyer de Schauensee 1970) placed in the monotypic genus Gnathospiza,
but Paynter (1970a) merged this into Sicalis, following the suggestion
by Meyer de Schauensee (1966); this treatment has been followed by most
subsequent authors (e.g., Ridgely & Tudor 1989, Ridgely & Greenfield
2001).
100.
Evidence for maintaining Emberizoides duidae as a separate species from E.
herbicola is weak (Hilty 2003); they were treated as conspecific by
Hellmayr (1938), Phelps & Phelps (1950a), and Paynter (1970a); <check
Eisenmann & Short 1982> ; they form a superspecies (Sibley & Monroe
1990).
101.
Eisenmann & Short (1982) showed that E. ypiranganus is widely
sympatric with E. herbicola and thus merits continued recognition as a
separate species.
102.
Genetic data indicate that Emberizoides and Embernagra belong in
the Thraupidae (Burns et al. 2002, 2003, Klicka et al. 2007, Campagna et al.
2011) and that Emberizoides and Embernagra are sister taxa
(Klicka et al. 2007). SACC proposal passed to move to Thraupidae.
103.
Sibley & Monroe (1990) considered Embernagra platensis and E.
longicauda to form a superspecies.
104.
Nores et al. (1983) reported sympatry between olivascens and nominate platensis
in C—rdoba, which would elevate the former to species rank; they were formerly
(e.g., REF) treated as separate species but considered conspecific by Meyer de
Schauensee (1966). <incorp. Contreras 1980>. Hayes (2003) provided
additional evidence for lack of gene flow between the olivascens group
and nominate platensis. SACC proposal to treat olivascens group as a
separate species did not pass.
105.
Meyer de Schauensee (1970) called this species "Buff-throated
Pampa-Finch", and Ridgely & Tudor (1989) called it "Pale-throated
Serra-Finch."
106.
Genetic data indicate that Volatinia belongs in the Thraupidae (Burns et
al. 2002, 2003, Klicka et al. 2007) and may be the sister genus to Conothraupis
(Burns et al. 2003). SACC proposal passed to move to Thraupidae.
Steadman (1982) proposed that Volatinia be merged with Geospiza,
and this was followed by Wetmore et al. (1984); however, nest
architecture and genetic data (Burns et al. 2002, Klicka et al. 2007) do not
support this.
107.
Clark (1986) found that Volatinia, Sporophila, Oryzoborus [now Sporophila], Dolospingus [now
Sporophila], and Charitospiza share a unique foot scute
character that suggests that they form a monophyletic group. Genetic data
(Klicka et al. 2007), however, indicate that Volatinia and Sporophila
+ Oryzoborus are not close relatives. Mason & Burns (2013) found that Dolospingus was embedded in Sporophila and proposed merger of it
into Sporophila. SACC proposal passed to merge Oryzoborus and Dolospingus into Sporophila. Dickinson & Christidis (2014)
followed this treatment.
108.
Genetic data (Lijtmaer et al. 2004, Campagna et al. 2009, 2013) indicate that
levels of genetic divergence among species in the genus Sporophila are
quite low compared to other genera and that some taxa currently treated as
species may be paraphyletic. In perhaps no other genus of Neotropical birds are
there so many species-level taxa of that are suspected to represent aberrant
individuals or hybrids. Traditionally, relationships within the
genus have based on plumage similarities, but these have been shown to be influenced
by convergence by Mason & Burns (2013), who found that only 1 of the 7
plumage groups of Ridgely & Tudor (1989) was monophyletic. SACC proposal passed to change linear sequence. A new species, Sporophila iberaensis,
was described by Di Giacomo and Kopuchian (2016). SACC proposal
badly needed.
109.
Genetic data (Bledsoe 1988, Burns et al. 2002, 2003, Klicka et al. 2007)
indicate that Sporophila and Oryzoborus belong in the Thraupidae;
some morphological data are consistent with this (Clark 1986). SACC proposal passed to move to Thraupidae. Olson
(1981c) provided evidence that there is little morphological evidence for
maintaining the genus Oryzoborus as separate from Sporophila, and
Wetmore et al. (1984) merged Oryzoborus into Sporophila; the
diagnosis of Oryzoborus relies on bill size and shape, which are
notoriously unreliable indicators of phylogenetic relationships, although see
Stiles (1996). Webster & Webster (1999) also recommended keeping them as
separate genera because of differences in skeletal morphology. Sick (1963)
noted that intergeneric hybrids were <numerous?>. Genetic data (Lijtmaer
et al. 2004) strongly support the merger of Oryzoborus into Sporophila
(and provide minimal support for monophyly of Oryzoborus itself). SACC proposal to merge Oryzoborus into Sporophila did not
pass. Mason & Burns (2013) found that Oryzoborus was embedded in Sporophila and proposed merger of it
into Sporophila. SACC proposal passed to merge Oryzoborus and Dolospingus into Sporophila.
110.
The genus Sporophila was formerly (e.g., REFS, Phelps & Phelps
1950a) known as Spermophila, but see <REF>.
111.
Genetic data (Lijtmaer et al. 2004, Mason & Burns 2013) indicate that Sporophila
falcirostris and S. schistacea are sister species, confirming their
traditional placement (e.g., Meyer de Schauensee 1970) in linear sequences.
111a. Repenning et al. (2013) described a new
species, Sporophila beltoni,
for the yellow-billed population of S. plumbea. SACC proposal passed to recognize Sporophila beltoni.
112.
Sporophila intermedia may include a cryptic species, S. insularis
(Rodner et al. 2000, Restall 2002, Hilty 2003). SACC proposal to recognize insularis as a
separate species did not pass.
113.
Sporophila corvina and S. americana are considered
conspecific by most recent authors (e.g., Meyer de Schauensee 1970, Paynter
1970a, Ridgely & Tudor 1989); Olson's (1981a) study of the contact zone in
Panama between corvina and the subspecies hicksii of S. americana
suggested complete intergradation between the two, with the taxon "aurita"
representing nothing more than a hybrid swarm. Subsequently, Stiles (1996b) provided
rationale for recognition of the corvina and americana groups as
separate species, representing a return to the classification of Hellmayr
(1938); Stiles (1996b) also treated the subspecies murallae as separate
species from S. americana, and this was followed by Dickinson
(2003). Stiles (1996b) also
provided rationale for treating S. intermedia as part of this complex. SACC proposal passed to revise species limits.
114.
Stiles (1996b) showed that the correct name for this species group is corvina,
not aurita as in many references. <check>
115.
Sporophila bouvronides was formerly (e.g., Meyer de Schauensee 1970,
Paynter 1970a) considered a subspecies of S. lineola, but see Schwartz
(1975) for rationale for treating them as separate species, representing a
return to the classification of Hellmayr (1938) and Phelps & Phelps
(1950a); this treatment has been followed by subsequent authors; they
constitute a superspecies (Sibley & Monroe 1990).
116.
"Sporophila melanops," known only from the type
specimen from Goi‡s, Brazil, and treated as a valid species by Hellmayr (1938)
and Pinto (1944), is usually treated as
species of uncertain status (Meyer de Schauensee 1970, Sibley & Monroe
1990); more likely a variant of S. nigricollis or a hybrid than a valid
species (Ridgely & Tudor 1989). Treated as a valid species by Dickinson et
al. (2003) but not by Dickinson % Christidis (2014). See Hybrids and Dubious Taxa.
117.
Sick (1997), based on Sick (1962, 1963), suggested that S. ardesiaca is
only a subspecies or variant of S. nigricollis; Meyer de Schauensee
(1966) and Ridgely & Tudor (1989) expressed doubt that ardesiaca was
a valid species. Dickinson & Christidis (2014) treated it as a subspecies
of S. nigricollis. <check Willis-Oniki 1993 reference therein>. SACC proposal badly needed.
118.
Genetic data (Lijtmaer et al. 2004, Campagna et al. 2009, 2013Mason & Burns
2013) indicate that Sporophila bouvreuil (including pileata), S. minuta, S. hypoxantha, S.
ruficollis, S. palustris, S. hypochroma, S. cinnamomea, S. nigrorufa, and
S. melanogaster form a monophyletic group, as reflected in traditional
linear sequences (e.g., Meyer de Schauensee 1970) based on plumage
similarities. These plumage similarities also suggest that if S. zelichi is a valid species (see Note 32a),
then it also belongs in this group (Ridgely & Tudor 1989). Lijtmaer et al. (2004) included S. castaneiventris in this group, but
see Campagna et al. (2009, 2013) and Mason & Burns (2013). SACC proposal
needed to change linear sequence.
119. The western subspecies bicolor was formerly (e.g.,
REF) treated as a separate species from Sporophila leucoptera; Ridgely & Tudor (1989) suggested that bicolor
might merit recognition as a separate species.
120.
Sporophila peruviana was formerly (e.g., Hellmayr 1938) placed in the
monotypic genus Neorhynchus.
121.
The subspecies saturata was formerly (e.g., Hellmayr 1938) considered a
separate species from Sporophila bouvreuil, but they were considered
conspecific by Meyer de Schauensee (1966). Machado & Silveira
(2011) showed that saturata is not a diagnosable taxon. They also demonstrated the subspecies pileata is sympatric with S. bouvreuil and therefore should be considered a
separate species. SACC proposal passed to
treat pileata as separate species.
Dickinson & Christidis
(2014) followed this treatment. SACC proposal passed to establish English names of S.
bouvreuil and S. pileata. Mason
& Burns (2013) found that S. bouvreuil and S. pileata might not be sister species. Campagna et al. (2013) found not only
that they were not sisters, but also that S. bouvreuil was likely the sister to all of the ÒcapuchinoÓ
group except for S. minuta.
122.
Ridgely & Tudor (1989) provided reasons for why Sporophila insulata
might be the sister species of S. telasco, rather than to S.
minuta, the traditional hypothesis. Sporophila telasco and S. minuta
were considered closely related, perhaps sister species, by Hellmayr (1938),
but subsequent classifications (e.g., Paynter 1970a) separated them without
explanation. Stiles (2004) concluded from plumage characters that S.
insulata is indeed more closely related to S. telasco and
also that it is more likely a color morph or subspecies of the latter. SACC proposal passed to delete Sporophila insulata as a valid
species. Dickinson & Christidis (2014)
followed this treatment.
123.
Sporophila hypoxantha was formerly (e.g., Hellmayr
1938, Pinto 1944, Meyer de Schauensee 1966, 1970) considered a
subspecies of S. minuta, but see Short (1969a); this treatment, a return
to the classification of REF, has been followed by subsequent authors (e.g.,
Paynter 1970a, Ridgely & Tudor 1989). Sibley & Monroe (1990) considered
them to form a superspecies, but genetic data (Lijtmaer et al 2004, Campagna et
al. 2009, 2013, Mason & Burns 2013) indicate that they are not sister
species.
124.
Short (1975) suggested that Sporophila ruficollis was not a valid
species but a color morph of S. [minuta] hypoxantha; they
have been traditionally considered closely related, perhaps sister species
(e.g., Hellmayr 1938), as confirmed by Campagna et al. (2013).
125.
Short (1975) considered Sporophila palustris to be just a color
morph of S. ruficollis, but syntopy has not yet been reported
(Ridgely & Tudor 1989); Hellmayr (1938) considered it more closely related
to S. hypoxantha.
126.
Hellmayr (1938) pointed out the close relationship between Sporophila hypochroma
and S. cinnamomea; Meyer de Schauensee (1966) suggested that they
might be conspecific; Sibley & Monroe (1990) considered S. hypochroma
and S. cinnamomea to form a superspecies but suggested that hypochroma
might just be a color morph of S. cinnamomea. Meyer de Schauensee
(1952) considered hypochroma to be a subspecies of S. castaneiventris,
but later treated it as a separate species (Meyer de Schauensee 1966), as
recently confirmed by genetic data (Lijtmaer et al. 2004, Campagna et al. 2009,
2013), which also indicate that they are not even sister species.
127.
Narosky (1977) described Sporophila zelichi as a new species, but
whether it is a valid species is controversial. Ridgely & Tudor (1989) and
Sibley & Monroe (1990) maintained it as a species but noted that it was
perhaps a localized color morph of S. cinnamomea or a hybrid
population (S. cinnamomea X S. palustris), as
suggested by Vuilleumier & Mayr (1987). Mazar Barnett & Pearman (2001)
also continued to recognize it as a species. The observations of Azpiroz (2003)
suggest that zelichi could be a valid species confined to marsh
vegetation. Areta (2008) presented evidence that there is no data to support
species rank for zelichi. SACC proposal passed to remove from main list.
128.
Hellmayr (1936) placed Oryzoborus with the carduelines, but
morphological data (Beecher REF, Tordoff 1954, Bock 1960) indicated that it was
emberizine, where moved by Meyer de Schauensee (1966).
129.
Olson (1981b,c) and most authors (e.g., Meyer de Schauensee 1970, Paynter 1970,
Ridgely & Tudor 1989, Ridgely & Greenfield 2001) have considered S.
funerea to be a subspecies of S. angolensis, with the composite
English name "Lesser Seed-Finch." However, the nature of gene flow
between the two has never been studied adequately; although hybrids occur in
the area of contact in northern Colombia, there is no evidence of a hybrid
swarm or intergradation (contra Ridgely & Greenfield 2001, Hilty 2003) in
the area that would indicate free interbreeding between the two; thus, AOU
(1983, 1998) continued to rank them as separate species, following Hellmayr
(1938). SACC proposal to consider S. funerea and S. angolensis
conspecific did not pass.
130.
Species limits in the large Oryzoborus [now Sporophila] seed-finches are complex, controversial,
and need further work. Paynter (1970a) treated them all as conspecific. Meyer
de Schauensee (1970x) provided rationale for why S. maximiliani
should be treated as a separate species from S. crassirostris;
they are broadly sympatric in the Guianas region; however, this sympatry might
be only during the nonbreeding season <>(Phelps & Phelps 1950).
Treatment as separate species has been followed by most subsequent authors
(e.g., Ridgely & Tudor 1989, REFS). However, as outlined by Ridgely &
Tudor (1989), placement of some subspecies is likely incorrect. Also, Sibley
& Monroe (1990), followed by Ridgely & Greenfield (2001), treated the
subspecies atrirostris (with gigantirostris) as a separate
species from S. maximiliani. Hellmayr (1938) treated atrirostris
(with gigantirostris) as a separate species. [get TSS to do
sentences on atrirostris]. Middle American S. nuttingi was
formerly (e.g., REFS) included in S. maximiliani, but see Stiles
(1984).
136.
Called "Large-billed Seed-finch" in Meyer de Schauensee (1970).
137.
Called "Greater Large-billed Seed-finch" in Meyer de Schauensee
(1970).
138.
Genetic data indicate that Catamenia belongs in the Thraupidae (Burns et
al. 2002, 2003, Klicka et al. 2007, Campagna et al. 2011) and forms a group
with Diglossa, Acanthidops, Xenodacnis, Diglossa, Haplospiza,
and most Phrygilus. SACC proposal passed to move to Thraupidae. Catamenia
was placed with the carduelines by Hellmayr (1938) but then moved to the
emberizines by Meyer de Schauensee (1966) based on the morphological data of
Beecher (REF) and Tordoff (1954).
139.
The Santa Marta subspecies oreophila was reluctantly treated as a
separate species by Hellmayr (1938), Meyer de Schauensee (1966), and Paynter
(1970a), but most recent authors, including Meyer de Schauensee (1970), have
treated it as a subspecies of C. homochroa; see Zambrano (1977)
for rationale.
140.
<>Miller (1928) proposed that Charitospiza was closely related to Lophospingus.
141.
Genetic data indicate that Coryphospingus belongs in the Thraupidae
(Burns et al. 2002, 2003, Klicka et al. 2007, Burns & Racicot 2009), as
suspected long ago by Paynter (1970a), and forms a group with Lanio, Eucometis,
and Tachyphonus. SACC proposal passed to move to Thraupidae.
142.
Coryphospingus pileatus and C. cucullatus form a superspecies
(Sibley & Monroe 1990); they hybridize to an uncertain extent in <>
(REF). Their sister relationship
was confirmed by Burns & Racicot (2009).
143.
Called "Gray Pileated-Finch" in Ridgely & Tudor (1989).
144.
Called "Red Pileated-Finch" in Ridgely & Tudor (1989) and Ridgely
& Greenfield (2001). Formerly
called "Crimson Finch", but this is the long-standing name for the
Old World estrildid Neochmia phaeton; see Clements and Shany (2001).
145.
Paynter (1971) suggested that Rhodospingus might belong in
Thraupidae. Burns & Racicot
(2009) found not only that it was a member of the Thraupidae but also that it
might be embedded within one group of Tachyphonus. SACC proposal passed to move to Thraupidae.
146.
The genus Gubernatrix has been placed traditionally in the Emberizidae,
sometimes (e.g., Hellmayr 1938, Meyer de
Schauensee 1966, 1970) with the cardinal grosbeaks, which in this
classification are considered a separate family, Cardinalidae. Paynter (1970a)
tentatively included it in the Emberizidae, following Tordoff (1954), who
presented morphological data that indicated that Gubernatrix was not
related to the cardinalines.
Genetic data indicate that it belongs in the Thraupidae (Campagna et al.
2011). SACC
proposal passed to move to Thraupidae.
147.
Until recently, the relationships of Coereba remained unresolved, and
temporary treatment as a monotypic family (e.g., AOU 1998) seemed warranted.
Some authors (Beecher 1951, Tordoff 1954a, Lowery & Monroe 1968) have
included it within the Parulidae, and others (e.g., Bledsoe 1988) have included
it within the Thraupidae; the most recent genetic data set (Burns et al. 2002)
provides strong support for a monophyletic group consisting of Coereba, Tiaris,
and the Galapagos finches (including Pinaroloxias), as well as Caribbean
genera Euneornis, Loxigilla, Loxipasser, Melanospiza,
Melopyrrha; this group appears to be embedded within the thraupine
lineage. Lovette & Bermingham's (2002) genetic data were also consistent
with placement of Coereba in the Thraupidae. SACC
proposal passed to abandon the family name
"Coerebidae" and to move Coereba elsewhere. SACC proposal passed to move next to Tiaris and
Galapagos "finches", and to place them all in Incertae Sedis
category.
148.
Genetic data indicate that Tiaris belongs in the Thraupidae (Burns et
al. 2002, 2003) as part of a group that includes Coereba, the Galapagos
finches, and several Caribbean genera (see also Sato et al. 2001, Yuri and
Mindell 2002, Klicka et al. 2007). SACC proposal passed move Tiaris and Galapagos
"finches" next to Coereba, and to place them all in Incertae
Sedis category. Tiaris itself is also paraphyletic, with minimally olivacea, the type
species of the genus, not being not particularly close to other "Tiaris."
Tiaris had formerly (e.g., Hellmayr 1938, AOU 1957) been placed with the
cardinalines, and then with the emberizines based on skeletal morphology
(Tordoff 1954a).
149.
Tiaris is masculine, so the correct spellings of the species names are olivaceus,
obscurus, and fuliginosus (David & Gosselin 2002b).
150.
Tiaris obscurus was formerly placed in the genus Sporophila and
known as "Dull-colored Seedeater" (e.g., Hellmayr 1938, Meyer de
Schauensee 1970), but nest structure and voice indicate that it belonged in Tiaris
s.l. (Paynter 1970a, Ridgely & Tudor 1989), as confirmed by recent genetic
(Burns et al. 2002, Lijtmaer et al. 2004) and morphological (Clark 1986)
data. Genetic data (Lijtmaer et al.
2004) indicate that Tiaris fuliginosus and T. obscurus are sister
species. The comprehensive
phylogenies of Burns et al. (2014) and Barker et al. (2015) showed that these
two species are not closely related to Tiaris; Burns et al. (2016) described a new genus, Asemospiza, for these two species. SACC proposal pending to recognize Asemospiza.
151. Based on the comprehensive phylogenies
of Burns et al. (2014) and Barker et al. (2015), Burns et al. (2016)
recommended the transfer of Tiaris bicolor
to Melanospiza (as in Dickinson &
Christidis 2014). SACC proposal pending.
152.
Genetic data indicate that the Galapagos finches belong in the Thraupidae
(Burns et al. 2002, 2003) as part of a group that includes Coereba, Tiaris,
and several Caribbean genera (see also Akie et al. 2001, Sato et al. 2001, Yuri
and Mindell 2002, Klicka et al. 2007). SACC proposal passed to move Tiaris and
Galapagos "finches" next to Coereba, and to place them all in
Incertae Sedis category. Classification of
Galapagos finches based on Petren et al. (1999), Sato et al. (1999, 2001), and
Burns et al. (2002), who found that resurrection of Platyspiza for crassirostris
is required to keep Camarhynchus from being paraphyletic (and that
extralimital Pinaroloxias is embedded within the Galapagos finches).
<incorp. Freeland and Boag 1999a.> The Galapagos finches were formerly
(e.g., Hellmayr 1938) placed in a separate subfamily, Geospizinae, from other
sparrows and finches, but were placed in the Emberizinae by Mayr and Amadon
(1951) and Tordoff (1954a). Subsequent genetic data (e.g., Petren et al. 1999,
Sato et al. 1999, 2001, Burns et al. 2002) have confirmed the monophyly of the
group.
153.
Genetic data (Petren et al. 1999, Sato et al. 1999, 2001, Burns et al. 2002)
indicate that Certhidea olivacea is basal within the Galapagos finches.
154.
Freeland & Boag (1999b), Petren et al. (1999), and Tonnis et al. (2005)
found that C. olivacea consists of two distinct lineages. Tonnis et al.
(2005) found that the two lineages were associated with habitat differences
among islands, with one (olivacea) found on islands with moist upland
woodland and the other (fusca) found on drier woodland on low islands;
the two lineages show no sings of reproductive isolation (Grant & Grant
2002). SACC proposal passed to recognize C. fusca as a separate species.
155.
"Camarhynchus conjunctus," known from two specimens
from Charles and treated as a valid species by Hellmayr (1938), is considered a
probable hybrid (Camarhynchus parvulus X Certhidea olivaceus)
(Lack 1947, Paynter 1970a). "Camarhynchus aureus," known
from one specimen from Chatham and treated as a valid species by Hellmayr
(1938), is also considered a probable hybrid (Camarhynchus parvulus X Certhidea
olivaceus) (Lack 1947, Paynter 1970a). "Camarhynchus giffordi,"
known from one specimen from Indefatigable and treated as a valid species
by Hellmayr (1938), is also considered an aberrant C. pallidus or hybrid
(Camarhynchus pallidus X Certhidea olivaceus) (Lack 1947, Paynter
1970a). See Hybrids and Dubious Taxa.
156.
Camarhynchus pallida and C. heliobates were formerly (e.g., Hellmayr 1938) treated in a separate genus, Cactospiza,
but REFS and Paynter (1970a) merged this into Camarhynchus; this merger is strongly supported by genetic data
(Freeland & Boag 1999b, Petren et al. 1999, Sato et al. 1999, 2001).
157.
The subspecies affinis and habeli were formerly (e.g., Hellmayr 1938) both considered a separate species
from Camarhynchus psittacula, but REFS and Paynter (1970a)
treated them as conspecific.
158.
Petren et al. (2005) showed that Geospiza difficilis might be
polyphyletic, with the central island populations perhaps sister to most of the
DarwinÕs Finches, although microsatellite and mtDNA differ in the pattern of
relationship of different populations.
Lamichhaney et al. (2015) proposed that difficilis consists of three species, with acutirostris
of Genovesa and septentrionalis of Wolf and Darwin elevated to species
rank. SACC
proposal passed to recognize three species. SACC
proposal pending on English names.
159.
See Paynter (1970a) over concerns that the name nebulosa may have
priority for this species.
160.
Large and small beak morphs of Geospiza fortis show positive assortative
mating on Santa Cruz Island (Huber et al. 2007).
161.
Petren et al. (2005) found that the Espa–ola population of Geospiza
conirostris may be distinct enough to deserve separation as a species. Lamichhaney et al. (2015) proposed that conirostris consists of two species, with propinqua
of Genovesa elevated to species rank.
SACC proposal passed to recognize three species. SACC
proposal pending on English names.
162.
Parkerthraustes humeralis was formerly (e.g., Hellmayr 1938, Pinto 1944, Meyer de Schauensee 1970, Paynter 1970c,
Ridgely & Tudor 1989, Sibley & Monroe 1990) placed in the genus Caryothraustes;
for rationale for generic separation of Parkerthraustes from Caryothraustes,
as anticipated by Hellmayr (1938), see Demastes & Remsen (1994) and Remsen
(1997). Recent genetic data (Klicka et al. 2007)
confirm that Parkerthraustes is not only not part of Caryothraustes
but also not a cardinaline and that it likely belongs in the Thraupidae. SACC proposal passed to transfer to Thraupidae.
163.
Saltator grossus and S. fuliginosus are sister allotaxa (Chaves et al. 2013) that might be best treated as
conspecific (Meyer de Schauensee 1966, Ridgely & Tudor 1989); evidence for
treatment of the two species as separate species is weak; they were treated as
conspecific by Paynter (1970c) and as forming a superspecies by Sibley &
Monroe (1990).
164.
Klicka et al. (2007) found strong genetic support for a sister relationship
between Saltator and core Thraupidae. Sushkin (1924) proposed that Saltator
was thraupine, not emberizine/cardinaline. SACC proposal passed to transfer Saltator from
Cardinalidae to Incertae Sedis. SACC proposal to transfer to Thraupidae did not pass.
Barker et al. (2013) found that Saltator
and Saltatricula were embedded in the
Thraupidae. SACC proposal passed to transfer to Thraupidae. Chavez
et al. (2013) found that relationships within the genus are not consistent with
the current linear sequence of species.
SACC proposal passed to revise linear sequence.
165.
Saltator grossus and S. fuliginosus were formerly placed in the
genus Pitylus (e.g., Hellmayr 1938, Pinto
1944, Meyer de Schauensee 1970, Paynter 1970c); for merger of Pitylus
into Saltator, see Hellack & Schnell (1977), Tamplin et al. (1993),
Demastes and Remsen (1994), Klicka et al. (2007), and
Chaves et al. (2013).
166.
Sibley & Monroe (1990) considered Saltator coerulescens, S.
similis, and S. maxillosus to form a superspecies, but see Note 11.
167.
Hilty (2003) treated the Middle American grandis subspecies group as a
separate species from the nominate South American Saltator coerulescens group,
a return to the classification of (REF). Hilty (2003) also indicated that vocal
differences within South America suggest that additional species may be involved. The gene tree of Chavez et al. (2013) is
consistent with treating the Middle American and northern South American grandis group as a separate species: if striatipectus is treated as a separate
species, it is the sister to the South American coerulescens group,
making broadly defined coerulescens paraphyletic. SACC proposal needed.
168.
Hellmayr (1938) considered Saltator maxillosus and S. aurantiirostris
conspecific, and Short (1975) reported that S. maxillosus and S.
aurantiirostris intergrade in Corrientes, Argentina <check>.
169.
Saltator nigriceps was formerly (e.g., Hellmayr 1938, Paynter
1970c, FjeldsŒ & Krabbe 1990) considered conspecific with S. aurantiirostris,
but most recent classifications have followed Meyer de Schauensee (1966) in
considering them separate species owing to differences in bill and tail shape
and body size. They are parapatric (Ridgely & Tudor 1989), and Sibley &
Monroe (1990) considered them to form a superspecies; they evidently differ in
vocalizations (Ridgely & Greenfield 2001). Klicka et al. (2007) and Chaves et al. (2013) found that nigriceps was actually basal to ((S. aurantiirostris + S. maxillosus)
+ (S. grossus + S. fuliginosus)).
170.
Saltator striatipectus was formerly (e.g., Hellmayr 1938, Meyer
de Schauensee 1970, Paynter 1970c) considered conspecific with S. albicollis
("Lesser Antillean Saltator"), but see Seutin et al. (1993) for a
return to the classification of (REF); Seutin et al. (1993) also suggested that
additional South American taxa might deserve recognition as separate
species. However, Chaves et al. (2013) found that Saltator striatipectus was indeed part of the S. albicollis/S.
coerulescens group. SACC proposal passed to revise linear sequence.
171.
Inclusion of S. cinctus and S. atricollis in Saltator was
questioned by Hellack and Schnell
(1977)(? REFs). However, Chaves et al. (2013) found that S. cinctus was deeply embedded in Saltator. See also Note
172.
172.
Saltatricula, traditionally placed in the Emberizidae, is one of many
"emberizine" genera for which genetic data (Burns et al. 2003)
suggest a closer relationship to the Thraupidae. Klicka et al. (2007) found
that it was embedded within the genus Saltator (and that Saltator
itself was close to if not sister to Thraupidae) and that its sister species
was Saltator atricollis. SACC proposal passed to transfer out of Emberizidae and
place next to Saltator. SACC proposal to merge into Saltator did not pass. Chaves et al. (2013)
confirmed the sister relationship between Saltator atricollis and
Saltatricula multicolor and found that
these two were the sister taxon to all other Saltator; therefore, Saltatricula should be
merged into Saltator, or atricollis should be transferred to Saltatricula.
SACC proposal passed to revise linear sequence. SACC proposal to merge Saltatricula into Saltator did not pass. Dickinson & Christidis (2014) moved atricollis
to Saltatricula. Burns et al. (2016) recommended that Saltatricula
be merged into Saltator. SACC proposal passed to place atricollis in Saltatricula.
INCERTAE SEDIS
[Almost certainly do not belong in their traditional
families, listed here Incertae Sedis; see Burns (1997) Burns et al. (2002,
2003), Klicka et al. (2000), Yuri & Mindell (2002).]
Mitrospingus cassinii Dusky-faced
Tanager 8a
Mitrospingus oleagineus Olive-backed
Tanager 8a
Rhodinocichla rosea Rosy
Thrush-Tanager 9, 9a
8a. Barker et al. (2013) found Mitrospingus is a member of a lineage
that is not particularly close to other nine-primaried families and proposed creating
a new family, Mitrospinigdae, for it, Orthogonys,
and Lamprospiza. SACC proposal
needed <wait on NACC>
9.
Familial affinities of Rhodinocichla rosea have
always been uncertain, with some suspecting that it might be closest to the
Mimidae (Skutch 1962), but traditionally placed in the tanagers (Eisenmann
1962), with support from morphological data (Clark 1974, Raikow 1978); genetic
data (Seutin and Bermingham 1997) suggest that it is closest to some
"tanagers". Storer (1970a) suspected that plumage similarities
between Rhodinocichla and Granatellus suggested a close
relationship between the two.
Barker et al. (2013) found that it was not particularly closely related
to any other nine-primaried oscine lineage and that to keep existing families
monophyletic, they proposed a monotypic family for it, Rhodinocichlidae; this
was adopted by Dickinson & Christidis (2014). SACC proposal
needed <wait on NACC>.
9a.
Formerly (e.g., Meyer de Schauensee 1970) known as "Rose-breasted
Thrush-Tanager."
EMBERIZIDAE (SPARROWS) 1
Oreothraupis arremonops Tanager Finch 49
Chlorospingus flavigularis Yellow-throated Chlorospingus
66
Chlorospingus parvirostris Short-billed Chlorospingus
66, 67
Chlorospingus canigularis Ashy-throated Chlorospingus
68
Chlorospingus flavopectus Common Chlorospingus
64, 65, 65a, 69
Chlorospingus tacarcunae Tacarcuna Chlorospingus
69
Chlorospingus semifuscus Dusky Chlorospingus
70
Chlorospingus flavovirens Yellow-green Chlorospingus
67a
Rhynchospiza stolzmanni Tumbes Sparrow 3
Rhynchospiza strigiceps Stripe-capped Sparrow 3
Ammodramus savannarum Grasshopper Sparrow 2
Ammodramus humeralis Grassland Sparrow 2
Ammodramus aurifrons Yellow-browed Sparrow 2
Arremonops conirostris Black-striped Sparrow 44b
Arremonops tocuyensis Tocuyo Sparrow 44, 44a
Arremon basilicus Sierra Nevada Brushfinch 47, 47a, 48, 48a
Arremon perijanus Perija Brushfinch 47, 48
Arremon atricapillus Black-headed Brushfinch 47, 48
Arremon phaeopleurus Caracas Brushfinch 47, 48
Arremon phygas Paria Brushfinch 47, 48
Arremon assimilis Gray-browed Brushfinch 47, 48
Arremon torquatus White-browed Brushfinch 47, 48
Arremon aurantiirostris Orange-billed Sparrow 45a
Arremon abeillei Black-capped Sparrow 45a, 45c
Arremon schlegeli Golden-winged Sparrow 45a
Arremon taciturnus Pectoral Sparrow 45, 45a, 45b, 47
Arremon franciscanus S‹o Francisco Sparrow 45a, 46
Arremon semitorquatus
Half-collared Sparrow 45
Arremon flavirostris Saffron-billed Sparrow 45a
Arremon brunneinucha Chestnut-capped Brushfinch 47
Arremon castaneiceps Olive Finch 47, 49a
Zonotrichia capensis Rufous-collared Sparrow
Melospiza lincolnii Lincoln's Sparrow (V)
1a
Atlapetes albinucha White-naped Brushfinch 47a, 50a
Atlapetes albofrenatus Moustached Brushfinch 50
Atlapetes personatus Tepui Brushfinch
Atlapetes melanocephalus Santa Marta Brushfinch
Atlapetes semirufus Ochre-breasted Brushfinch
Atlapetes flaviceps Yellow-headed Brushfinch 51
Atlapetes fuscoolivaceus Dusky-headed Brushfinch
Atlapetes leucopis White-rimmed Brushfinch
Atlapetes albiceps White-headed Brushfinch 54c
Atlapetes rufigenis Rufous-eared Brushfinch 53
Atlapetes tricolor Tricolored Brushfinch 51a
Atlapetes schistaceus Slaty Brushfinch 54a, 55a
Atlapetes pallidinucha Pale-naped Brushfinch
Atlapetes blancae Antioquia Brushfinch 52c
Atlapetes latinuchus Yellow-breasted Brushfinch 52, 52a, 52b
Atlapetes leucopterus White-winged Brushfinch 54b
Atlapetes pallidiceps Pale-headed Brushfinch 54c
Atlapetes seebohmi Bay-crowned Brushfinch 55, 55a, 55b
Atlapetes nationi Rusty-bellied Brushfinch 55, 55a
Atlapetes forbesi Apurimac Brushfinch 53
Atlapetes melanopsis Black-spectacled Brushfinch 54
Atlapetes terborghi Vilcabamba Brushfinch 57, 52
Atlapetes canigenis Cuzco Brushfinch 56, 54a
Atlapetes melanolaemus Black-faced Brushfinch 58, 58a, 52
Atlapetes rufinucha Bolivian Brushfinch 52, 52aa
Atlapetes fulviceps Fulvous-headed Brushfinch
Atlapetes citrinellus Yellow-striped Brushfinch
1.
Genetic data (Bledsoe 1988, Sibley & Ahlquist 1990, Lougheed et al. 2000,
Burns et al. 2002, 2003, Klicka et al. 2007, Sedano & Burns 2010 -- check
Groth-Barrowclough etc.) indicate that the family Emberizidae as traditionally
constituted is polyphyletic, with most genera occurring in South America
belonging to the tanager lineage; some morphological data (Clark 1986) also
support this. The only genera in South America
traditionally placed in the Emberizidae for which genetic data indicate that
they are true Emberizidae are: Zonotrichia, Ammodramus, Aimophila
(DaCosta et al. 2009), Arremon, and Atlapetes; the majority have
been found to be members of the Thraupidae; see Note 1 under that family. Barker et al. (2013) and Klicka et al.
(2014) found that even a more narrowly defined Emberizidae was not a
monophyletic group and that recognition of a new family, Passerellidae, was
required for all New World members of Emberizidae. SACC
proposal needed <wait on NACC> Klicka
et al. (2014) also found that the phylogenetic relationships among genera and
species in this family are not reflected in traditional linear sequences. SACC proposal passed to modify linear sequence.
1a.
Recorded from Aruba (Voous 1985).
2.
Ammodramus humeralis and A. aurifrons were formerly
(e.g., Hellmayr 1938, Pinto 1944, Phelps &
Phelps 1950a, Meyer de Schauensee 1970) treated in a separate genus, Myospiza,
but most recent authors (e.g., Ridgely & Tudor 1989, Sibley and Monroe
1990) have followed Paynter (1970a) and Robins & Schnell (1971) in merging
this into Ammodramus. Genetic data (Carson & Spicer 2003, Klicka
& Spellman 2007, DaCosta et al. 2009, Klicka et al. 2014) indicate that as
currently defined, Ammodramus is polyphyletic. Because the type species
of Ammodramus is savannarum, and DaCosta et al. (2009) and Klicka
et al. (2014) found that the latter is the sister to the two Myospiza,
this result should not affect classification of South American species.
3. Rhynchospiza
was traditionally (e.g., Hellmayr 1938, Meyer
de Schauensee 1970) treated as a monotypic genus, with its sole species being stolzmanni; however, most recent authors
(e.g., Ridgely & Tudor 1989) have followed Paynter (1967, 1970a) in merging
this into Aimophila, which was widely suspected of being polyphyletic
(Ridgway 1901, Storer 1955b, Wolf 1977).
DaCosta et al. (2009) have confirmed that Aimophila is polyphyletic and that the South American taxa are not
members of true Aimophila; they
recommended resurrection of Rhynchospiza,
which would also include stolzmanniÕs
sister species, A. strigiceps. SACC proposal passed to resurrect Rhynchospiza. Klicka et al. (2014) found that Rhynchospiza is sister to the group of
sparrow genera that includes true Ammodramus,
Arremonops, and Peucaea.
44.
The genus Arremonops has been merged into Arremon by some authors
(Phelps and Phelps 1950a, Meyer de Schauensee 1951), but they are not closely
related; see Klicka et al. (2014).
44a.
Arremonops tocuyensis was considered to form a superspecies with Middle
American A. rufivirgatus (Mayr & Short 1970, AOU 1983, Sibley
and Monroe 1990), but it is the sister species to A. conirostris (Klicka et al. 2014). Arremonops conirostris was
formerly (e.g., Hellmayr 1938) considered conspecific with Middle American A.
chloronotus, but they are sympatric in Honduras (Monroe 1963b), and Klicka
et al. (2014) found that A. chloronotus is sister to A. rufivirgatus.
45.
Raposo and Parrini (1997) proposed recognizing the subspecies semitorquatus
as a separate species from Arremon taciturnus. SACC
proposal passed to recognize semitorquatus
as a species.
45a.
Sibley and Monroe (1990) considered all the species of Arremon form a
superspecies, but A. aurantiirostris and A. abeillei are
sympatric in western Ecuador.
45b.
The Colombian subspecies axillaris was treated as a separate species from Arremon taciturnus by
<REF>.
45c.
The Mara–on subspecies nigriceps was treated as
a separate species from Arremon abeillei by <REF>.
46.
Described since Meyer de Schauensee (1970): Raposo (1997).
47.
Buarremon was merged into Atlapetes by Hellmayr (1938), and this
was followed by Paynter (REFS, 1970a), Meyer de Schauensee (1966, 1970), and
most subsequent authors. Remsen and Graves (1995) resurrected the genus Buarremon
as separate from Atlapetes because it is not certain that they are
sister genera; this treatment was followed by AOU (1998), Ridgely et al.
(2001), and Dickinson (2003). Genetic data (DaCosta et al. 2009, Klicka et al.
2014) confirm that Buarremon is not
closely related to Atlapetes. Cadena et al. (2007) found that Buarremon
itself is paraphyletic with respect to Arremon and probably Lysurus. SACC proposal passed to merge all into Arremon; also followed by Banks et al. (2008). Additional genetic analyses
(Fl—rez-Rodr’guez et al. 2011) found that the paraphyly of Buarremon was
generated largely by mtDNA gene trees and that some, but not all, other loci
support a monophyletic Buarremon. Klicka et
al. (2014) confirmed that former Buarremon
was indeed paraphyletic with respect to Arremon
and Lysurus.
47a.
The placement of the former Buarremon
in Arremon creates a problem with the
English names in that hyphenated ÒBrush-FinchÓ now used in two genera, Atlapetes and Arremon, with most species in the latter called ÒSparrowÓ or
ÒFinch.Ó Thus, the hyphen is
misleading with respect to relationships, and this needs fixing. SACC proposal passed to change to ÒBrushfinchÓ.
48.
The relationships among the forms assigned to the atricapillus and torquatus
groups are controversial, with virtually no relevant data available. Wetmore et al. (1984), Paynter (1970a),
and Remsen & Graves (1995) treated the atricapillus group as
conspecific with B. torquatus largely because of the intermediate
phenotypes shown by subspecies such as tacarcunae and costaricensis.
Hellmayr (1938), Meyer de Schauensee (1966), Sibley & Monroe (1990), and
Ridgely & Tudor (1989) treated them as two species because of the close
geographical approach of nominate atricapillus and B. t. assimilis
without signs of gene flow. Donegan
et al. (2007) found B. [t.] atricapillus and B. [t.]
assimilis to replace one another elevationally in the East Andes of
Colombia (Santander and Boyac‡ departments), without any evidence of
hybridization, suggesting that treatment of this complex within a single
species is not supportable. Sibley
& Monroe (1990) considered B. torquatus and B. atricapillus,
along with Middle American B. virenticeps, to form a superspecies. Buarremon
virenticeps was considered conspecific with B. torquatus by Paynter
(1970a) and Wetmore et al. (1984), but was treated as a separate species by
Paynter (1978), AOU (1983, 1998), and Ridgely & Tudor (1989). Cadena & CuervoÕs (2010) analysis of
voice, plumage, and genetics in the group indicates that as many as eight
species should be recognized in this complex. SACC proposal passed to revise species limits.
48a.
Called ÒBangsÕs Brush-finchÓ in Hilty (2011) and Dickinson & Christidis
(2014).
49.
The relationships of Oreothraupis have been controversial. It was included in the Thraupidae by
Hellmayr (1936). Storer (1958)
pointed out that similarities in plumage pattern and texture, and in juvenal
plumage strongly suggested a close relationship to Atlapetes, and
Paynter (1970a) considered it close enough to Atlapetes that he stated
that they might be considered congeneric.
Klicka et al. (2014) found that it was a New World sparrow
(Passerellidae) and that it was sister to Chlorospingus.
49a.
Sibley & Monroe (1990) considered A.
castaneiceps to form a superspecies with Middle American A.
crassirostris. Genetic
data (Klicka et al. 2014) confirm that they are sister species, as anticipated
from their former treatment as the two species in the genus Lysurus. See Note 47.
50.
Donegan & Huertas (2006) found no strong support in an analysis of plumage
characters for A. a. meridae (MŽrida range) being closely related to the
nominate race (Eastern Cordillera) and that the two taxa differ in biometrics;
however, vocal and molecular analyses are lacking.
50a.
Paynter (1964) provided rationale for merging the Atlapetes gutturalis
group into A. albinucha; and this treatment was followed by Paynter
(1970a), AOU (1998), and Dickinson (2003).
51.
Formerly (e.g., Meyer de Schauensee 1970, Hilty & Brown 1986, Dickinson
2003) known as "Olive-headed Brush-Finch". Called "Yellow-headed
Brush-Finch" in BirdLife International (2000). SACC proposal passed to change English name.
51a. The subspecies crassus of the W. Andes of Colombia and Ecuador may merit species
rank (Ridgely & Greenfield 2001).
Dickinson (2003) prematurely elevated crassus to species rank.
52.
Atlapetes rufinucha was formerly (Hellmayr
1938, Paynter 1970a, Meyer de Schauensee 1970, Ridgely & Tudor 1989,
Sibley & Monroe 1990) considered to be a polytypic species with a disjunct
distribution. However, the genetic
data of Garc’a-Moreno and FjeldsŒ (1999) and Klicka et al. (2014) corroborated
the predictions of Remsen & Graves (1995b) that these populations did not
form a monophyletic group but instead were more closely related to adjacent
populations of A. schistaceus. Thus, Atlapetes latinuchus was
formerly considered a subspecies of A. rufinucha, but it is more
closely related to parapatric A. pallidinucha (Klicka et al. 2014).
Donegan & Huertas (2006) noted that A. latinuchus itself
(even without A. [l.] nigrifrons) might involve more than
one species. See also Note 54a
below.
52a.
Called "Northern Rufous-naped Brush-Finch" in Garc’a-Moreno and
FjeldsŒ (1999) and "Cloud-forest Brush-Finch" in Clements and Shany
(2001). SACC proposal to change English name did not pass.
52aa. Broadly defined Atlapetes rufinucha was called ÒRufous-naped Brush-Finch,Ó and so
narrowly defined A. rufinucha was
renamed ÒBolivian Brush-FinchÓ to avoid confusion. SACC proposal passed to change English name.
52b.
Donegan & Huertas (2006) and Donegan et al. (2014) proposed that the
subspecies nigrifrons (formerly phelpsi) of the Perij‡ Mountains
should be ranked as a species from Atlapetes
latinuchus. SACC proposal needed.
52c.
Donegan (2007b) described a new species (Atlapetes blancae) that is
possibly most closely related to the A. latinuchus group, from the
Central Andes of Colombia. SACC proposal passed to recognize A. blancae.
53.
Atlapetes forbesi has traditionally been treated (e.g., Meyer de
Schauensee 1970, Paynter 1970c, Ridgely & Tudor 1989) as a subspecies of
A. rufigenis, although Meyer de Schauensee (1966) suggested that it should
be treated as a separate species. Genetic data (Garc’a-Moreno & FjeldsŒ
1999, Klicka et al. 2014) indicate that they are each more closely related to
other species of Atlapetes than they are to each other and thus must be
treated as separate species.
54.
Described since Meyer de Schauensee (1970): Valqui & FjeldsŒ (1999). More
recently renamed: Valqui & FjeldsŒ (2002).
54a.
Atlapetes schistaceus was formerly (Hellmayr
1938, Paynter 1970a, Meyer de Schauensee 1970, Ridgely & Tudor 1989,
Sibley & Monroe 1990) considered to be a polytypic species with a disjunct
distribution. However, the genetic data of Garc’a-Moreno and FjeldsŒ (1999) and
Klicka et al. (2014) corroborated the predictions of Remsen & Graves
(1995b) that these populations did not form a monophyletic group but instead
were more closely related to parapatric populations of A. rufinucha.
Thus, Atlapetes canigenis was formerly considered a subspecies of A.
schistaceus, but it is more closely related to parapatric members of the rufinucha
group.
54b.
[paynteri etc.] The subspecies dresseri
was formerly (e.g., REF) considered a separate species from Atlapetes
leucopterus, but they were treated as conspecific by Paynter (REF) and
Meyer de Schauensee (1966).
54c.
Hellmayr (1938) and Meyer de Schauensee (1966) suggested that A. pallidiceps
should be considered a subspecies of A. albiceps, but they are not
sister taxa (Klicka et al. 2014).
55.
For continued treatment of A. seebohmi and A. nationi
as separate species, as in Hellmayr (1938) and Meyer de Schauensee (1970), see
Ridgely & Tudor (1989); Koepcke (1957, 1958), Paynter (1970a, 1972), and
FjeldsŒ & Krabbe (1990) regarded them as conspecific; they form a
superspecies. Paynter (1970a)
suspected that A. nationi (with seebohmi) might be better treated
as a subspecies of A. schistaceus.
55a.
Sibley & Monroe (1990) considered Atlapetes schistaceus, A. seebohmi,
and A. nationi to form a superspecies; however, A. schistaceus
is a paraphyletic species (see Note 54a), and so if this superspecies
designation is correct, it applies only to the nominate schistaceus
group and must also include A. latinuchus. Klicka et al. (2014) found that
A. latinuchus and A.
pallidinucha.
55b.
The subspecies ceciliae and simonsi were formerly (e.g., Hellmayr 1938) considered separate species from Atlapetes
seebohmi, but Paynter (1970a) and Meyer de Schauensee (1966) treated them
as conspecific.
56.
Called "Grey Brush-Finch" in Garc’a-Moreno and FjeldsŒ (1999) and
"Cusco Brush-Finch" in Clements and Shany (2001). SACC proposal passed to change English name from
"Sooty Brush-Finch," as in Dickinson (2003), to "Cuzco
Brush-Finch".
57.
Recently described (as a subspecies of A. rufinucha): Remsen (1993).
Garc’a-Moreno & FjeldsŒ (1995) provided evidence that it should be recognized
as a separate species. SACC proposal to lump terborghi into melanolaemus
or rufinucha did not pass.
58.
Atlapetes melanolaemus was formerly (Hellmayr
1938, Paynter 1970a, Meyer de Schauensee 1970, Ridgely & Tudor 1989,
Sibley & Monroe 1990) considered a subspecies of A. rufinucha, but
see Garc’a-Moreno & FjeldsŒ (1999). SACC proposal to lump melanolaemus into rufinucha
did not pass.
58a.
Called "Dark-faced Brush-Finch" in Clements and Shany (1999).
64.
Genetic data (REFS, Burns et al. 2002, 2003) indicate the genus Chlorospingus
is not a member of the Thraupidae, but (Klicka et al. 2007) a member of the
Emberizidae. SACC proposal passed to transfer to Emberizidae. Barker
et al. (2013) further confirmed the placement of the genus with the New World
sparrows. Frank Pitelka (in Tordoff
1954a) long ago noted the emberizine-like behavior of Chlorospingus. SACC proposal passed to change English names of the
species in the genus from ÒBush-TanagerÓ to ÒChlorospingusÓ. Called
ÒTanager FinchesÓ by Dickinson & Christidis (2014).
65.
Chlorospingus ophthalmicus, as currently circumscribed, is paraphyletic
with respect to C. tacarcunae, C. semifuscus, and C. inornatus,
and likely consists of several species (Weir et al. 2008).
65a. Zimmer (1947) evidently did not realize
that the name flavopectus Lafresnaye, 1840, has priority over ophthalmicus (Du Bus de
Gisignies, 1847). SACC proposal
pending to use flavopectus. SACC
proposal passed to use flavopectus.
66.
Chlorospingus parvirostris was formerly (e.g., Hellmayr 1936) considered
a subspecies of C. flavigularis, but see Zimmer (1947).
67.
Chlorospingus parvirostris was called "Yellow-whiskered Bush-Tanager"
in Isler & Isler (1987), Ridgely & Tudor (1989), Sibley & Monroe
(1990), and Ridgely et al. (2001). Proposal needed.
67a.
Klicka et al. (2014) found that flavovirens
was not a member of Chlorospingus and
was actually a true tanager (Thraupidae).
Avenda–o et al. (2016) found that its closest relatives were Bangsia tanagers and recommend its
transfer to that genus. SACC proposal badly needed.
68.
Stiles & Skutch (1989) suggested that the isolated Central American
subspecies, olivaceiceps, may deserve recognition as a separate species
from South American Chlorospingus canigularis.
68a.
Recent genetic data (Yuri & Mindell 2002, Burns et al. 2003) failed to find
support for inclusion of Mitrospingus in Thraupidae. Klicka et al.
(2007) found that it did not fit within any of the traditionally recognized
families.
69.
Species limits in the Chlorospingus ophthalmicus complex are
controversial. Sibley & Monroe (1990) considered Chlorospingus
ophthalmicus and C. tacarcunae to form a superspecies with C.
inornatus of Cerro Pirre, eastern Panama; C. tacarcunae was formerly
(e.g., Hellmayr 1936) considered a subspecies of C. flavigularis before
Zimmer (1947) treated it as a subspecies of C. ophthalmicus. Meyer de
Schauensee (1966, 1970) continued to treat tacarcunae as a subspecies of
C. ophthalmicus, but most classifications have followed Ridgely (1976),
AOU (1983), and Wetmore et al. (1984) in treating it as a separate species. The
subspecies cinereocephalus of central Peru was formerly (e.g., Hellmayr
1936) considered a subspecies of C. semifuscus until Zimmer (1947)
considered it a subspecies of C. ophthalmicus. The flavopectus
subspecies group of Ecuador and northern Peru was formerly (e.g., Hellmayr 1936) treated as a separate species from C.
ophthalmicus, but see Zimmer (1947).
70.
Formerly (e.g., Meyer de Schauensee 1970, FjeldsŒ & Krabbe 1990) known as
"Dusky-bellied Bush-Tanager."
Part 11. Oscine Passeriformes, C (Cardinalidae to end)